সংরক্ষণ করুন:
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| বিন্যাস: | Recurso digital |
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Zenodo
2019
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| বিষয়গুলি: | |
| অনলাইন ব্যবহার করুন: | https://doi.org/10.5281/zenodo.10447785 |
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সূচিপত্রের সারণি:
- <p><b><i>Landouria madurensis</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: 024A8B78-91C2-49CA-8721-C0B8CCC46982</p><p>Figs 8, 22, 36, 53, 57; Tables 3–4</p><p><i>Helix squamulosa</i> Martens, 1867: 266, pl. 13, fig. 10 (“ Insel Madura (an der Nordostseite von Java), bei Sumanap”) (objective junior synonym of <i>Helix squamulosa</i> Mousson, 1856). <b>Syn. nov.</b></p><p><i>Helix squamulosa</i> – Pfeiffer 1871 in 1870–1876: pl. 117, figs 13–15; 1872 in 1870–1876: 41. <i>Landouria squamulosa</i> – van Benthem Jutting 1941: 311.</p><p><b>Diagnosis</b></p><p><i>Landouria madurensis</i> sp. nov. is characterized by a sharply keeled shell with longish scales all over the teleoconch, a wide umbilicus, a long penis, which is divided into a longer, broader distal and a shorter, narrower proximal part, and a long flagellum, which gradually tapers towards its proximal end.</p><p><b>Etymology</b></p><p><i>Landouria madurensis</i> sp. nov. is named after its occurrence on Madura Island.</p><p><b>Material examined</b></p><p><b>Holotype</b></p><p>INDONESIA • holotype (det. anat.); East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest; 6°54′01″ S, 113°12′08″ E; 17 m a.s.l.; 14 May 2014; A.S. Nurinsiyah and T. Setiadi leg.; D = 11.4 mm, H = 6.5 mm; MZB 21432.</p><p><b>Paratypes</b></p><p>INDONESIA • 1 spec.; same data as for holotype; MZB 19209 • 4 spec.; same data as for holotype; MZB 19210 • 1 spec.; same data as for holotype; ZMH 133408 • 4 spec.; same data as for holotype; ZMH 133437 • 2 spec. (also syntypes of <i>H. squamulosa</i>); Sumenep (see Martens 1867: 267); 7°01′ S, 113°52′ E; ZMB 7155.</p><p><b>Description</b></p><p>SHELL (Figs 8, 57; Tables 3–4). Depressed conical, with 5.5–5.75 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; without distinct incised spiral lines; longish scales all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, very wide, comprising 38–40% of shell diameter, hardly obscured by columellar edge.</p><p>GENITALIA (Figs 22, 36; Table 4). Atrium short; penis long, divided into a longer, broader distal and a shorter, narrower proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum long, broadest in its middle part, gradually tapering towards its proximal end, slightly crenated, without node; vas deferens narrow and long; vagina cylindrical, long; oviduct shorter, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.</p><p><b>Remarks</b></p><p><i>Landouria madurensis</i> sp. nov. differs from the neighbouring <i>Landouria intumescens</i> in the presence of longish scales all over the teleoconch (Fig. 8), a wider umbilicus (U/D 0.38–0.40 vs 0.22–0.34 in <i>L. intumescens</i>) and a relatively long vagina (P:V 2.1 vs 2.4–3.9 in <i>L. intumescens</i>).</p><p>A second, more conical species of <i>Landouria</i> without longish scales and with a narrower umbilicus, of which only juveniles were found, lives sympatrically with <i>Landouria madurensis</i> sp. nov. in Nepa Monkey lowland rainforest on Madura Island.</p><p><b>Distribution</b></p><p><i>Landouria madurensis</i> sp. nov. is endemic to Madura Island (Fig. 53).</p>