I tiakina i:
Ngā taipitopito rārangi puna kōrero
Ngā kaituhi matua: Zhang, Le-Jia, Shu, Shu-Sen, Song, Xin-Yuan, Naing, Nay Htet, Oo, Thaung Naing, Chen, Xiao-Yong
Hōputu: Recurso digital
Reo:
I whakaputaina: Zenodo 2025
Ngā marau:
Urunga tuihono:https://doi.org/10.5281/zenodo.15080299
Ngā Tūtohu: Tāpirihia he Tūtohu
Kāore He Tūtohu, Me noho koe te mea tuatahi ki te tūtohu i tēnei pūkete!
Rārangi ihirangi:
  • <p><i>Hydrobioides physcus</i> Annandale, 1918 comb. rev.</p><p><i>Hydrobioides physcus</i> Annandale, 1918: 121–122, pl. 13, figs 8, 8 a, 9, pl. 14, figs 5, 5 a (Myanmar, Shan State, Inle Lake).</p><p><i>Hydrobioides</i> (<i>Paranerita</i>) <i>physcus</i> – Annandale, 1920: 45.</p><p><i>Paranerita physcus</i> – Annandale & Rao, 1925: 115.</p><p><i>Parabithynia physcus</i> – Pilsbry, 1928: 108</p><p><i>Gabbia physcus</i> – Subba Rao, 1989: 78, figs 127–129.</p><p><b>Diagnosis.</b></p><p>Shell medium, low spiral whorls, weak keel on shoulder, outer lip outward extended.</p><p><b>Material examined.</b></p><p>• 3 probable <i>paratypes</i> (preserved dry), Inle Lake, South Shan, Burma. “ Ex. India Museum ”, NHMUK.20060144; 2 specimens (preserved in ethanol), Shan State, Taunggyi District, Inn Paw Hkon, Inthein, riverbank under Inthein Bridge (Fig. 1 B. d), 20°27'35.9"N, 96°50'32.6"E, 1 Jul. 2024, Le-Jia Zhang leg., KIZ.2400061 –2400062 • 2 specimens (preserved in ethanol), Shan State, Taunggyi District, Nyaungshwe, Inle Lake, floating islands near Intha (Fig. 1 B. c), 20°36'33.4"N, 96°54'53.8"E, 3 Jul. 2024, Le-Jia Zhang leg., KIZ.2400063 –2400064.</p><p><b>Description.</b></p><p>Shell (Fig. 7) medium for the family (Table 1), relatively thick, solid, subglobose to globose, white or bright yellow to orange yellow, with four whorls at adulthood; teleoconch low, with a keel on shoulder, forming weak nodules on shoulder with vertical growth lines, two weak additional keels on body whorl; aperture ovate, lip thickened, always more than half of shell in height, outer lip outward extended; umbilicus covered by inner lip completely.</p><p>Operculum (Figs 7 A, 4 B) calcium, ovate, thin, slightly smaller than aperture, transparent grey; exterior surface smooth, nucleus close to the central of operculum; interior surface with relatively thin outer opercular region, narrow weak concentric veins on the margin of inner opercular region, nuclear region relatively large, with irregular veins and grains.</p><p>Penis (Fig. 5 B) with a white penial appendix located in the central of penis, penial appendix slightly shorter than distal part of penis.</p><p>Radula (Fig. 8) taenioglossate; central tooth upper margin with one broad triangle central cusp and two to three small sharp cusps on either side, lateral margin each with two to three small sharp cusps; lateral tooth with one broad tongue-shape central cusp and two to three small sharp cusps; inner marginal tooth with 10 to 11 small cusps; outer marginal tooth with six small cusps.</p><p><b>Remarks.</b></p><p>This species can be easily distinguished from other bithyniid snails based on the thick subglobose to globose shell with keels and shoulder. It also can be distinguished from <i>H. nassa</i> based on opercular characters and less small cusps on inner and outer marginal teeth. This species is the type species and only extant species of genus <i>Parabithynia</i> Pilsbry, 1928, which is now endemic to Inle Lake. Several fossil species of this genus were reported from China and Russia (Popova 1981; Yu et al. 1978), but the assignment needs a revision. Our molecular study confirms that this species is the sister species of <i>H. nassa</i>, with a close genetic relationship (p distance of 16 S: 4.3–4.5 %). The similarity in shell (especially the extended outer lip), operculum, and male genital structure also supports that these two species should be placed in one genus. Therefore, we consider that <i>Parabithynia</i> is a synonym of <i>Hydrobioides</i>. <i>Hydrobioides physcus</i> comb. rev. should be re-assigned to its original genus.</p><p><b>Habitat and distribution.</b></p><p>Inle Lake and the big rivers connected to Inle Lake; swamps in Heho, Shan State, Myanmar.</p>