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Zenodo
2025
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| Online Access: | https://doi.org/10.5281/zenodo.19137121 |
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Table of Contents:
- <p><b><i>Bhatia</i> Distant, 1908</b></p><p><i>Bhatia</i> Distant, 1908: 357; Webb, 1994: 116; Zhang & Webb, 1996: 12; Viraktamath & Murthy, 2009: 47; Shang, Shen, Zhang & Li, 2006: 565; Lu & Zhang, 2015: 146; Yu, Qu, Dai & Yang, 2019: 143; Lu & Zhang, 2024: 278.</p><p><b>Type species:</b> <i>Eutettix? olivacea</i> Melichar, 1903.</p><p><b>Diagnosis.</b> Head short, crown broadly rounded anteriorly, with transverse carina; median length slightly greater than length next to eye; transverse groove near anterior margin; head slightly wider than pronotum. Ocelli marginal, distant from eye. Antennae long, inserted at upper angle of eye. Hind femur apical setal formula 2+2+1.</p><p>Male pygofer with numerous macrosetae on lateral lobe, with or without an inner marginal ridge; if present, the ridge runs obliquely from inner margin to ventroposterior corner. Subgenital plate nearly triangular, inner margin nearly straight. Valve nearly quadrate. Connective Y-shaped or H-shaped; in some species, with a sclerotized plate or an elongate process between connective and aedeagus. Aedeagus with one or more pairs of basal processes.</p><p><b>Remarks.</b> Species of <i>Bhatia</i> are similar in external morphology and male genitalia, but can be immediately separated from <i>Bhatiahamus</i> by the absence of a pair of stout apical processes on the male tenth abdominal segment. Furthermore, this genus is also similar to <i>Fistulatus</i>, from which it differs most notably in lacking posterior elongations on the pygofer lobe.</p><p><b>Distribution.</b> Oriental Region and Pacific areas.</p><p><b>Checklist of Chinese species of <i>Bhatia</i></b></p><p><i>B</i>. <i>biconjugara</i> Zhang & Zhang, 1998</p><p><i>B</i>. <i>biconjugara</i> Zhang & Zhang, 1998: 178; Shang, Shen, Zhang & Li, 2006: 566; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Sichuan, Guangxi, Guangdong, Fujiang).</p><p><i>B. bilateritra</i> <b>sp. nov.</b></p><p>Distribution: China (Hainan).</p><p><i>B</i>. <i>digitata</i> Shang & Shen, 2006</p><p><i>B</i>. <i>digitata</i> Shang & Shen, 2006: 568; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Anhui, Henan, Guangxi).</p><p><i>B</i>. <i>furcata</i> <b>sp. nov.</b></p><p>Distribution: China (Yunnan).</p><p><i>B</i>. <i>hastata</i> Shang & Shen, 2006</p><p><i>B</i>. <i>hastata</i> Shang & Shen, 2006: 567; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Guangxi).</p><p><i>B. holodonta</i> <b>sp. nov.</b></p><p>Distribution: China (Yunnan).</p><p><i>B</i>. <i>koreana</i> (Kwon & Lee, 1979)</p><p><i>Koreanopsis koreana</i> Kwon & Lee, 1979: 50.</p><p><i>B</i>. <i>koreana</i> Zhang & Webb, 1996: 12.</p><p>Distribution: China (Guangxi, Shaanxi); South Korea.</p><p><i>B</i>. <i>affinikoreana</i> <b>sp. nov.</b></p><p>Distribution: China (Hainan).</p><p><i>B</i>. <i>loba</i> Lu & Zhang, 2024</p><p><i>B</i>. <i>loba</i> Lu & Zhang, 2024: 283.</p><p>Distribution: China (Tibet).</p><p><i>B</i>. <i>multispinosa</i> Lu & Zhang, 2015</p><p><i>B</i>. <i>multispinosa</i> Lu & Zhang, 2015: 148; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Sichuan).</p><p><i>B</i>. <i>olivacea</i> (Melichar, 1903)</p><p><i>Eutettix? olivaceus</i> Melichar, 1903: 191.</p><p><i>B</i>. <i>olivacea</i> (Melichar, 1903), Viraktamath, 1998: 171; Yu, Qu, Dai & Yang, 2019: 143.</p><p><i>Melichariella olivacea</i> Matsumura, 1914: 237.</p><p><i>B</i>. <i>divacea</i> Evans, 1947: 230.</p><p>Distribution: China (Hainan); India; Sri Lanka; Thailand; Java; Indonesia.</p><p><i>B</i>. <i>parasagittata</i> Lu & Zhang, 2024</p><p><i>B</i>. <i>parasagittata</i> Lu & Zhang, 2024: 281.</p><p>Distribution: China (Guizhou).</p><p><i>B</i>. <i>quadrispinosa</i> Shang & Zhang, 2006</p><p><i>B</i>. <i>quadrispinosa</i> Shang & Zhang, 2006: 571; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Sichuan).</p><p><i>B</i>. <i>sagittata</i> Cai & Shen, 1999</p><p><i>B</i>. <i>sagittata</i> Cai & Shen, 1999: 38.</p><p>Distribution: China (Henan, Jiangxi).</p><p><i>B</i>. <i>satsumensis</i> (Matsumura, 1914)</p><p><i>Melichariella satsumensis</i> Matsumura, 1914: 237.</p><p><i>B</i>. <i>satsumensis</i> Esaki & Ito, 1954: 94: Zhang & Webb, 1996: 12; Zhang & Zhang, 1998: 179; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Guangdong); Japan.</p><p><i>B</i>. <i>unicornis</i> Shang & Li, 2006</p><p><i>B</i>. <i>unicornis</i> Shang & Li, 2006: 573; Yu, Qu, Dai & Yang, 2019: 143.</p><p>Distribution: China (Guangdong).</p><p><b>Key to Chinese species of <i>Bhatia</i> (males)</b></p><p>1 Aedeagus with single basal process.............................................. <i>B. unicornis</i> Shang & Li, 2006</p><p>– Aedeagus with two or more basal processes................................................................ 2</p><p>2 Aedeagal shaft with processes........................................................................... 3</p><p>– Aedeagal shaft without processes........................................................................ 8</p><p>3 Aedeagal shaft with denticulate lateral flanges.............................................................. 4</p><p>– Aedeagal shaft without denticulate lateral flanges........................................................... 5</p><p>4 Style apex rounded, subapex acute (Fig. 48)............................................... <i>B. holodonta</i> <b>sp. nov.</b></p><p>– Style apex acute, subapex rounded................................................ <i>B. koreana</i> Kwon & Li, 1979</p><p>5 Aedeagal shaft with two pairs of processes (Figs. 13, 19).......................... <i>B. satsumensis</i> (Matsumura, 1914)</p><p>– Aedeagal shaft with only one pair of processes............................................................. 6</p><p>6 Processes at apex of aedeagal shaft arrow-shaped in ventral view............................................... 7</p><p>– Processes at apex of aedeagal shaft not above (Figs. 4, 10).............................. <i>B. olivacea</i> (Melichar, 1903)</p><p>7 Aedeagal shaft gradually tapered subapically (Figs. 18, 24)........................ <i>B. parasagittata</i> Lu & Zhang, 2024</p><p>– Aedeagal shaft distinctly inflated subapically (Figs. 3, 9).............................. <i>B. sagittata</i> Cai & Shen, 1999</p><p>8 Aedeagus with one pair of basal processes................................................................. 9</p><p>– Aedeagus with two or more pairs of basal processes........................................................ 14</p><p>9 Leaf-shaped sclerite articulated between aedeagus and connective (Figs. 6, 12)............... <i>B. loba</i> Lu & Zhang, 2024</p><p>– Leaf-shaped sclerite articulated absent between aedeagus and connective........................................ 10</p><p>10 Aedeagal shaft with basal processes crossing ventrally (Figs. 41–42).............................. <i>B. furcata</i> <b>sp. nov.</b></p><p>– Aedeagal shaft with basal processes not crossing ventrally................................................... 11</p><p>11 Aedeagal shaft medially inflated, not inflated apically, in ventral view (Fig. 32).................. <i>B. bilateritra</i> <b>sp. nov.</b></p><p>– Aedeagal shaft apically inflated, not inflated medially, in ventral view (Fig. 22).................................. 12</p><p>12 Style with apical process long and straight........................................ <i>B. hastata</i> Shang & Shen, 2006</p><p>– Style with apical process short and hooklike.............................................................. 13</p><p>13 Margins of basal aedeagal processes smooth (Fig. 22)............................... <i>B. digitata</i> Shang & Shen, 2006</p><p>– Margins of basal aedeagal processes undulate (Fig. 60).................................... <i>B. affinikoreana</i> <b>sp. nov.</b></p><p>14 Style apex rounded, subapex acute......................................... <i>B. biconjugara</i> Zhang & Zhang, 1998</p><p>– Style apex acute, subapex rounded...................................................................... 15</p><p>15 Single lateral process present between aedeagus and connective (Fig. 23)............. <i>B. multispinosa</i> Lu & Zhang, 2015</p><p>– No process between aedeagus and connective (Fig. 7)......................... <i>B. quadrispinosa</i> Shang & Zhang, 2006</p>