A general multi-scale description of metastable adaptive motion across fitness valleys

Fuente: arXiv
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Autores principales: Esser, Manuel, Kraut, Anna
Formato: Preprint
Publicado: 2021
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author Esser, Manuel
Kraut, Anna
author_facet Esser, Manuel
Kraut, Anna
contents We consider a stochastic individual-based model of adaptive dynamics on a finite trait graph $G=(V,E)$. The evolution is driven by a linear birth rate, a density dependent logistic death rate an the possibility of mutations along the (possibly directed) edges in $E$. We study the limit of small mutation rates for a simultaneously diverging population size. Closing the gap between the works of Bovier, Coquille and Smadi (2019) and Coquille, Kraut and Smadi (2021), we give a precise description of transitions between evolutionary stable conditions (ESC), where multiple mutations are needed to cross a valley in the fitness landscape. The system shows a metastable behaviour on several divergent time scales associated to a degree of stability. We develop the framework of a meta graph that is constituted of ESCs and possible metastable transitions between those. This allows for a concise description of the multi-scale jump chain arising from concatenating several jumps. Finally, for each of the various time scale, we prove the convergence of the population process to a Markov jump process visiting only ESCs of sufficiently high stability.
format Preprint
id arxiv_https___arxiv_org_abs_2112_12675
institution arXiv
publishDate 2021
record_format arxiv
spellingShingle A general multi-scale description of metastable adaptive motion across fitness valleys
Esser, Manuel
Kraut, Anna
Probability
Populations and Evolution
37N25, 60J27, 60J80, 92D15, 92D25
We consider a stochastic individual-based model of adaptive dynamics on a finite trait graph $G=(V,E)$. The evolution is driven by a linear birth rate, a density dependent logistic death rate an the possibility of mutations along the (possibly directed) edges in $E$. We study the limit of small mutation rates for a simultaneously diverging population size. Closing the gap between the works of Bovier, Coquille and Smadi (2019) and Coquille, Kraut and Smadi (2021), we give a precise description of transitions between evolutionary stable conditions (ESC), where multiple mutations are needed to cross a valley in the fitness landscape. The system shows a metastable behaviour on several divergent time scales associated to a degree of stability. We develop the framework of a meta graph that is constituted of ESCs and possible metastable transitions between those. This allows for a concise description of the multi-scale jump chain arising from concatenating several jumps. Finally, for each of the various time scale, we prove the convergence of the population process to a Markov jump process visiting only ESCs of sufficiently high stability.
title A general multi-scale description of metastable adaptive motion across fitness valleys
topic Probability
Populations and Evolution
37N25, 60J27, 60J80, 92D15, 92D25
url https://arxiv.org/abs/2112.12675