Error-induced extinction in a multi-type critical birth-death process

Fuente: arXiv
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Autori principali: Guasch, Meritxell Brunet, Krapivsky, P. L., Antal, Tibor
Natura: Preprint
Pubblicazione: 2023
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author Guasch, Meritxell Brunet
Krapivsky, P. L.
Antal, Tibor
author_facet Guasch, Meritxell Brunet
Krapivsky, P. L.
Antal, Tibor
contents Extreme mutation rates in microbes and cancer cells can result in error-induced extinction (EEX), where every descendant cell eventually acquires a lethal mutation. In this work, we investigate critical birth-death processes with $n$ distinct types as a birth-death model of EEX in a growing population. Each type-$i$ cell divides independently $(i)\to(i)+(i)$ or mutates $(i)\to(i+1)$ at the same rate. The total number of cells grows exponentially as a Yule process until a cell of type-$n$ appears, which cell type can only die at rate one. This makes the whole process critical and hence after the exponentially growing phase eventually all cells die with probability one. We present large-time asymptotic results for the general $n$-type critical birth-death process. We find that the mass function of the number of cells of type-$k$ has algebraic and stationary tail $(\text{size})^{-1-χ_k}$, with $χ_k=2^{1-k}$, for $k=2,\dots,n$, in sharp contrast to the exponential tail of the first type. The same exponents describe the tail of the asymptotic survival probability $(\text{time})^{-χ_n}$. We present applications of the results for studying extinction due to intolerable mutation rates in biological populations.
format Preprint
id arxiv_https___arxiv_org_abs_2306_11609
institution arXiv
publishDate 2023
record_format arxiv
spellingShingle Error-induced extinction in a multi-type critical birth-death process
Guasch, Meritxell Brunet
Krapivsky, P. L.
Antal, Tibor
Populations and Evolution
Probability
Extreme mutation rates in microbes and cancer cells can result in error-induced extinction (EEX), where every descendant cell eventually acquires a lethal mutation. In this work, we investigate critical birth-death processes with $n$ distinct types as a birth-death model of EEX in a growing population. Each type-$i$ cell divides independently $(i)\to(i)+(i)$ or mutates $(i)\to(i+1)$ at the same rate. The total number of cells grows exponentially as a Yule process until a cell of type-$n$ appears, which cell type can only die at rate one. This makes the whole process critical and hence after the exponentially growing phase eventually all cells die with probability one. We present large-time asymptotic results for the general $n$-type critical birth-death process. We find that the mass function of the number of cells of type-$k$ has algebraic and stationary tail $(\text{size})^{-1-χ_k}$, with $χ_k=2^{1-k}$, for $k=2,\dots,n$, in sharp contrast to the exponential tail of the first type. The same exponents describe the tail of the asymptotic survival probability $(\text{time})^{-χ_n}$. We present applications of the results for studying extinction due to intolerable mutation rates in biological populations.
title Error-induced extinction in a multi-type critical birth-death process
topic Populations and Evolution
Probability
url https://arxiv.org/abs/2306.11609