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Hauptverfasser: Carignano, Marcelo, Kröger, Martin, Almassalha, Luay, Agrawal, Vasundhara, Li, Wing Shun, Pujadas, Emily M., Nap, Rikkert J., Backman, Vadim, Szleifer, Igal
Format: Preprint
Veröffentlicht: 2023
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Online-Zugang:https://arxiv.org/abs/2310.02257
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author Carignano, Marcelo
Kröger, Martin
Almassalha, Luay
Agrawal, Vasundhara
Li, Wing Shun
Pujadas, Emily M.
Nap, Rikkert J.
Backman, Vadim
Szleifer, Igal
author_facet Carignano, Marcelo
Kröger, Martin
Almassalha, Luay
Agrawal, Vasundhara
Li, Wing Shun
Pujadas, Emily M.
Nap, Rikkert J.
Backman, Vadim
Szleifer, Igal
contents We propose the Self Returning Excluded Volume (SR-EV) model for the structure of chromatin based on stochastic rules and physical interactions. The SR-EV rules of return generate conformationally-defined domains observed by single cell imaging techniques. From nucleosome to chromosome scales, the model captures the overall chromatin organization as a corrugated system, with dense and dilute regions alternating in a manner that resembles the mixing of two disordered bi-continuous phases. This particular organizational topology is a consequence of the multiplicity of interactions and processes occurring in the nuclei, and mimicked by the proposed return rules. Single configuration properties and ensemble averages show a robust agreement between theoretical and experimental results including chromatin volume concentration, contact probability, packing domain identification and size characterization, and packing scaling behavior. Model and experimental results suggest that there is an inherent chromatin organization regardless of the cell character and resistant to an external forcing such as Rad21 degradation.
format Preprint
id arxiv_https___arxiv_org_abs_2310_02257
institution arXiv
publishDate 2023
record_format arxiv
spellingShingle Local Volume Concentration, Packing Domains and Scaling Properties of Chromatin
Carignano, Marcelo
Kröger, Martin
Almassalha, Luay
Agrawal, Vasundhara
Li, Wing Shun
Pujadas, Emily M.
Nap, Rikkert J.
Backman, Vadim
Szleifer, Igal
Biological Physics
Computational Physics
We propose the Self Returning Excluded Volume (SR-EV) model for the structure of chromatin based on stochastic rules and physical interactions. The SR-EV rules of return generate conformationally-defined domains observed by single cell imaging techniques. From nucleosome to chromosome scales, the model captures the overall chromatin organization as a corrugated system, with dense and dilute regions alternating in a manner that resembles the mixing of two disordered bi-continuous phases. This particular organizational topology is a consequence of the multiplicity of interactions and processes occurring in the nuclei, and mimicked by the proposed return rules. Single configuration properties and ensemble averages show a robust agreement between theoretical and experimental results including chromatin volume concentration, contact probability, packing domain identification and size characterization, and packing scaling behavior. Model and experimental results suggest that there is an inherent chromatin organization regardless of the cell character and resistant to an external forcing such as Rad21 degradation.
title Local Volume Concentration, Packing Domains and Scaling Properties of Chromatin
topic Biological Physics
Computational Physics
url https://arxiv.org/abs/2310.02257