Diffusion-driven self-assembly of emerin nanodomains at the nuclear envelope

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Main Authors: Alas, Carlos D., Wu, Liying, Pinaud, Fabien, Haselwandter, Christoph A.
Format: Preprint
Published: 2024
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author Alas, Carlos D.
Wu, Liying
Pinaud, Fabien
Haselwandter, Christoph A.
author_facet Alas, Carlos D.
Wu, Liying
Pinaud, Fabien
Haselwandter, Christoph A.
contents Emerin, a nuclear membrane protein with important biological roles in mechanotransduction and nuclear shape adaptation, self-assembles into nanometer-size domains at the inner nuclear membrane. The size and emerin occupancy of these nanodomains change with applied mechanical stress as well as under emerin mutations associated with Emery-Dreifuss muscular dystrophy (EDMD). Through a combination of theory and experiment we show here that a simple reaction-diffusion model explains the self-assembly of emerin nanodomains. Our model yields quantitative agreement with experimental observations on the size and occupancy of emerin nanodomains for wild-type emerin and EDMD-associated mutations of emerin, with and without applied forces, and allows successful prediction of emerin diffusion coefficients from observations on the overall properties of emerin nanodomains. Our results provide a physical understanding of EDMD-associated defects in emerin organization in terms of changes in key reaction and diffusion properties of emerin and its nuclear binding partners.
format Preprint
id arxiv_https___arxiv_org_abs_2407_11758
institution arXiv
publishDate 2024
record_format arxiv
spellingShingle Diffusion-driven self-assembly of emerin nanodomains at the nuclear envelope
Alas, Carlos D.
Wu, Liying
Pinaud, Fabien
Haselwandter, Christoph A.
Biological Physics
Soft Condensed Matter
Statistical Mechanics
Adaptation and Self-Organizing Systems
Subcellular Processes
Emerin, a nuclear membrane protein with important biological roles in mechanotransduction and nuclear shape adaptation, self-assembles into nanometer-size domains at the inner nuclear membrane. The size and emerin occupancy of these nanodomains change with applied mechanical stress as well as under emerin mutations associated with Emery-Dreifuss muscular dystrophy (EDMD). Through a combination of theory and experiment we show here that a simple reaction-diffusion model explains the self-assembly of emerin nanodomains. Our model yields quantitative agreement with experimental observations on the size and occupancy of emerin nanodomains for wild-type emerin and EDMD-associated mutations of emerin, with and without applied forces, and allows successful prediction of emerin diffusion coefficients from observations on the overall properties of emerin nanodomains. Our results provide a physical understanding of EDMD-associated defects in emerin organization in terms of changes in key reaction and diffusion properties of emerin and its nuclear binding partners.
title Diffusion-driven self-assembly of emerin nanodomains at the nuclear envelope
topic Biological Physics
Soft Condensed Matter
Statistical Mechanics
Adaptation and Self-Organizing Systems
Subcellular Processes
url https://arxiv.org/abs/2407.11758