HEIST: A Graph Foundation Model for Spatial Transcriptomics and Proteomics Data

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Main Authors: Madhu, Hiren, Rocha, João Felipe, Huang, Tinglin, Viswanath, Siddharth, Krishnaswamy, Smita, Ying, Rex
Format: Preprint
Published: 2025
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author Madhu, Hiren
Rocha, João Felipe
Huang, Tinglin
Viswanath, Siddharth
Krishnaswamy, Smita
Ying, Rex
author_facet Madhu, Hiren
Rocha, João Felipe
Huang, Tinglin
Viswanath, Siddharth
Krishnaswamy, Smita
Ying, Rex
contents Single-cell transcriptomics and proteomics have become a great source for data-driven insights into biology, enabling the use of advanced deep learning methods to understand cellular heterogeneity and gene expression at the single-cell level. With the advent of spatial-omics data, we have the promise of characterizing cells within their tissue context as it provides both spatial coordinates and intra-cellular transcriptional or protein counts. Proteomics offers a complementary view by directly measuring proteins, which are the primary effectors of cellular function and key therapeutic targets. However, existing models either ignore the spatial information or the complex genetic and proteomic programs within cells. Thus they cannot infer how cell internal regulation adapts to microenvironmental cues. Furthermore, these models often utilize fixed gene vocabularies, hindering their generalizability unseen genes. In this paper, we introduce HEIST, a hierarchical graph transformer foundation model for spatial transcriptomics and proteomics. HEIST models tissues as hierarchical graphs. The higher level graph is a spatial cell graph, and each cell in turn, is represented by its lower level gene co-expression network graph. HEIST achieves this by performing both intra-level and cross-level message passing to utilize the hierarchy in its embeddings and can thus generalize to novel datatypes including spatial proteomics without retraining. HEIST is pretrained on 22.3M cells from 124 tissues across 15 organs using spatially-aware contrastive and masked autoencoding objectives. Unsupervised analysis of HEIST embeddings reveals spatially informed subpopulations missed by prior models. Downstream evaluations demonstrate generalizability to proteomics data and state-of-the-art performance in clinical outcome prediction, cell type annotation, and gene imputation across multiple technologies.
format Preprint
id arxiv_https___arxiv_org_abs_2506_11152
institution arXiv
publishDate 2025
record_format arxiv
spellingShingle HEIST: A Graph Foundation Model for Spatial Transcriptomics and Proteomics Data
Madhu, Hiren
Rocha, João Felipe
Huang, Tinglin
Viswanath, Siddharth
Krishnaswamy, Smita
Ying, Rex
Genomics
Machine Learning
Cell Behavior
Single-cell transcriptomics and proteomics have become a great source for data-driven insights into biology, enabling the use of advanced deep learning methods to understand cellular heterogeneity and gene expression at the single-cell level. With the advent of spatial-omics data, we have the promise of characterizing cells within their tissue context as it provides both spatial coordinates and intra-cellular transcriptional or protein counts. Proteomics offers a complementary view by directly measuring proteins, which are the primary effectors of cellular function and key therapeutic targets. However, existing models either ignore the spatial information or the complex genetic and proteomic programs within cells. Thus they cannot infer how cell internal regulation adapts to microenvironmental cues. Furthermore, these models often utilize fixed gene vocabularies, hindering their generalizability unseen genes. In this paper, we introduce HEIST, a hierarchical graph transformer foundation model for spatial transcriptomics and proteomics. HEIST models tissues as hierarchical graphs. The higher level graph is a spatial cell graph, and each cell in turn, is represented by its lower level gene co-expression network graph. HEIST achieves this by performing both intra-level and cross-level message passing to utilize the hierarchy in its embeddings and can thus generalize to novel datatypes including spatial proteomics without retraining. HEIST is pretrained on 22.3M cells from 124 tissues across 15 organs using spatially-aware contrastive and masked autoencoding objectives. Unsupervised analysis of HEIST embeddings reveals spatially informed subpopulations missed by prior models. Downstream evaluations demonstrate generalizability to proteomics data and state-of-the-art performance in clinical outcome prediction, cell type annotation, and gene imputation across multiple technologies.
title HEIST: A Graph Foundation Model for Spatial Transcriptomics and Proteomics Data
topic Genomics
Machine Learning
Cell Behavior
url https://arxiv.org/abs/2506.11152