From shallow to full wrapping: geometry and deformability dictate lipid vesicle internalization

Fuente: arXiv
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Main Authors: van der Ham, Stijn, Brown, Alexander, Kusumaatmaja, Halim, Vutukuri, Hanumantha Rao
Format: Preprint
Published: 2025
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author van der Ham, Stijn
Brown, Alexander
Kusumaatmaja, Halim
Vutukuri, Hanumantha Rao
author_facet van der Ham, Stijn
Brown, Alexander
Kusumaatmaja, Halim
Vutukuri, Hanumantha Rao
contents The deformability of vesicles critically influences their engulfment by lipid membranes, a process central to endocytosis, viral entry, drug delivery, and intercellular transport. While theoretical models have long predicted this influence, direct experimental validation has remained elusive. Here, we combine experiments with continuum simulations to quantify how vesicle deformability affects the engulfment of small giant unilamellar vesicles (GUVs) by larger GUVs under depletion-induced adhesion. Using 3D confocal reconstructions, we extract vesicle shape, curvature, wrapping fraction, and the bendo-capillary length, a characteristic length scale that balances membrane bending and adhesion forces. We find that when vesicle size exceeds this length scale, engulfment is primarily governed by geometry. In contrast, when vesicle size is comparable to this scale, deformability strongly affects the transition between shallow, deep, and fully wrapped states, leading to suppression of full engulfment of vesicles. These findings connect theoretical predictions with direct measurements and offer a unified framework for understanding vesicle-mediated uptake across both synthetic and biological systems, including viral entry, synthetic cell design, drug delivery, and nanoparticle internalization.
format Preprint
id arxiv_https___arxiv_org_abs_2507_17434
institution arXiv
publishDate 2025
record_format arxiv
spellingShingle From shallow to full wrapping: geometry and deformability dictate lipid vesicle internalization
van der Ham, Stijn
Brown, Alexander
Kusumaatmaja, Halim
Vutukuri, Hanumantha Rao
Soft Condensed Matter
Biological Physics
The deformability of vesicles critically influences their engulfment by lipid membranes, a process central to endocytosis, viral entry, drug delivery, and intercellular transport. While theoretical models have long predicted this influence, direct experimental validation has remained elusive. Here, we combine experiments with continuum simulations to quantify how vesicle deformability affects the engulfment of small giant unilamellar vesicles (GUVs) by larger GUVs under depletion-induced adhesion. Using 3D confocal reconstructions, we extract vesicle shape, curvature, wrapping fraction, and the bendo-capillary length, a characteristic length scale that balances membrane bending and adhesion forces. We find that when vesicle size exceeds this length scale, engulfment is primarily governed by geometry. In contrast, when vesicle size is comparable to this scale, deformability strongly affects the transition between shallow, deep, and fully wrapped states, leading to suppression of full engulfment of vesicles. These findings connect theoretical predictions with direct measurements and offer a unified framework for understanding vesicle-mediated uptake across both synthetic and biological systems, including viral entry, synthetic cell design, drug delivery, and nanoparticle internalization.
title From shallow to full wrapping: geometry and deformability dictate lipid vesicle internalization
topic Soft Condensed Matter
Biological Physics
url https://arxiv.org/abs/2507.17434