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Autores principales: Galpern, Ezequiel A., Bueno, Carlos, Sánchez, Ignacio E., Wolynes, Peter G., Ferreiro, Diego U.
Formato: Preprint
Publicado: 2025
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Acceso en línea:https://arxiv.org/abs/2508.08109
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author Galpern, Ezequiel A.
Bueno, Carlos
Sánchez, Ignacio E.
Wolynes, Peter G.
Ferreiro, Diego U.
author_facet Galpern, Ezequiel A.
Bueno, Carlos
Sánchez, Ignacio E.
Wolynes, Peter G.
Ferreiro, Diego U.
contents We show how to localize and quantify the functional evolutionary constraints on natural proteins. The method compares the perturbations caused by local sequence variants to the energetics of the protein folding process and to the corresponding change to the apparent selection landscape of sequences over the evolutionary time scale. The difference between the physical folding free energies and the evolutionary free energies can be called a "Dark Energy". We analyse various protein sets and thereby show that Dark Energy is largely localized at functional sites, which are often energetically frustrated from the point of view of folding. Overall, we find that about 25% of the positions of the folded globular proteins display some significant Dark Energy. When a function relies on a free energy that can be thermodynamically quantified, such as a binding energy to a partner, the relationship of this physical free energy with Dark Energy can be used to define a Functional Selection Temperature. We show that selection for folding and binding functions bear similar weights in specific protein-protein interactions.
format Preprint
id arxiv_https___arxiv_org_abs_2508_08109
institution arXiv
publishDate 2025
record_format arxiv
spellingShingle Probing the Dark Energy in the Functional Protein Universe
Galpern, Ezequiel A.
Bueno, Carlos
Sánchez, Ignacio E.
Wolynes, Peter G.
Ferreiro, Diego U.
Biomolecules
We show how to localize and quantify the functional evolutionary constraints on natural proteins. The method compares the perturbations caused by local sequence variants to the energetics of the protein folding process and to the corresponding change to the apparent selection landscape of sequences over the evolutionary time scale. The difference between the physical folding free energies and the evolutionary free energies can be called a "Dark Energy". We analyse various protein sets and thereby show that Dark Energy is largely localized at functional sites, which are often energetically frustrated from the point of view of folding. Overall, we find that about 25% of the positions of the folded globular proteins display some significant Dark Energy. When a function relies on a free energy that can be thermodynamically quantified, such as a binding energy to a partner, the relationship of this physical free energy with Dark Energy can be used to define a Functional Selection Temperature. We show that selection for folding and binding functions bear similar weights in specific protein-protein interactions.
title Probing the Dark Energy in the Functional Protein Universe
topic Biomolecules
url https://arxiv.org/abs/2508.08109