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Main Authors: Nunns, Henry, Riehl, Manda
Format: Preprint
Published: 2025
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Online Access:https://arxiv.org/abs/2512.15828
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author Nunns, Henry
Riehl, Manda
author_facet Nunns, Henry
Riehl, Manda
contents In this paper we present two modifications of traditional $NK$ fitness landscapes, the $θNK$ and $HNK$ models, and explore these modifications via accessibility and ruggedness. The $θNK$ model introduces a parameter $θ$ to integrate local Rough Mount Fuji-type correlations in subgenotype contributions, simulating more biologically realistic correlated fitness effects. The $HNK$ model incorporates gene regulation effects by introducing a masking mechanism where certain loci modulate the expression of other loci, simulating effects observed in gene regulatory networks without modeling the full network. Through extensive simulations across a wide range of parameters ($N$, $K$, $θ$, and $H$), we analyze the impact of these modifications on landscape accessibility and the number of local optima. We find that increasing $θ$ or the number of masking loci ($H$) generally enhances accessibility, even in landscapes with many local optima, showing that ruggedness doesn't necessarily hinder evolutionary pathways. Additionally, distinct interaction patterns (blocked, adjacent, random) lead to different observations in accessibility and optimum structure. While more complex than traditional $NK$, we believe each model provides a new biologically relevant facet to fitness landscapes and provides insight into how genetic and regulatory structures influence the evolutionary potential of populations.
format Preprint
id arxiv_https___arxiv_org_abs_2512_15828
institution arXiv
publishDate 2025
record_format arxiv
spellingShingle Accessibility of Modified NK Fitness Landscapes
Nunns, Henry
Riehl, Manda
Quantitative Methods
92-10, 06B99
In this paper we present two modifications of traditional $NK$ fitness landscapes, the $θNK$ and $HNK$ models, and explore these modifications via accessibility and ruggedness. The $θNK$ model introduces a parameter $θ$ to integrate local Rough Mount Fuji-type correlations in subgenotype contributions, simulating more biologically realistic correlated fitness effects. The $HNK$ model incorporates gene regulation effects by introducing a masking mechanism where certain loci modulate the expression of other loci, simulating effects observed in gene regulatory networks without modeling the full network. Through extensive simulations across a wide range of parameters ($N$, $K$, $θ$, and $H$), we analyze the impact of these modifications on landscape accessibility and the number of local optima. We find that increasing $θ$ or the number of masking loci ($H$) generally enhances accessibility, even in landscapes with many local optima, showing that ruggedness doesn't necessarily hinder evolutionary pathways. Additionally, distinct interaction patterns (blocked, adjacent, random) lead to different observations in accessibility and optimum structure. While more complex than traditional $NK$, we believe each model provides a new biologically relevant facet to fitness landscapes and provides insight into how genetic and regulatory structures influence the evolutionary potential of populations.
title Accessibility of Modified NK Fitness Landscapes
topic Quantitative Methods
92-10, 06B99
url https://arxiv.org/abs/2512.15828