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Main Authors: Staub, Davide, Rabe, Finn, Misra, Akhil, Pauli, Yves, Hüppi, Roya, Yang, Ni, Lang, Nils, Michels, Lars, Edkins, Victoria, Frühholz, Sascha, Sommer, Iris, Hinzen, Wolfram, Homan, Philipp
Format: Preprint
Published: 2025
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Online Access:https://arxiv.org/abs/2512.20481
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author Staub, Davide
Rabe, Finn
Misra, Akhil
Pauli, Yves
Hüppi, Roya
Yang, Ni
Lang, Nils
Michels, Lars
Edkins, Victoria
Frühholz, Sascha
Sommer, Iris
Hinzen, Wolfram
Homan, Philipp
author_facet Staub, Davide
Rabe, Finn
Misra, Akhil
Pauli, Yves
Hüppi, Roya
Yang, Ni
Lang, Nils
Michels, Lars
Edkins, Victoria
Frühholz, Sascha
Sommer, Iris
Hinzen, Wolfram
Homan, Philipp
contents To maintain coherence in language, the brain must satisfy key competing temporal demands: the gradual accumulation of meaning across extended context (drift) and the rapid reconfiguration of representations at event boundaries (shift). How these processes are implemented in the human brain during naturalistic listening remains unclear. Here, we tested whether both can be captured by annotation-free drift and shift signals and whether their neural expression shows distinct regional preferences across the brain. These signals were derived from a large language model (LLM) processing the narrative input. To enable high-precision voxelwise encoding models with stable parameter estimates, we densely sampled one healthy adult across more than 7 hours of listening to crime stories while collecting 7 Tesla fMRI data. We then modeled the feature-informed hemodynamic response using a regularized encoding framework validated on independent stories. Drift predictions were prevalent in default-mode network hubs, whereas shift predictions were evident bilaterally in the primary auditory cortex and language association cortex. Together, these findings show that coherence during language comprehension is implemented through distinct but co-expressed neural regimes of slow contextual integration and rapid event-driven reconfiguration, offering a mechanistic entry point for understanding disturbances of language coherence in psychiatric disorders.
format Preprint
id arxiv_https___arxiv_org_abs_2512_20481
institution arXiv
publishDate 2025
record_format arxiv
spellingShingle Coherence in the brain unfolds across separable temporal regimes
Staub, Davide
Rabe, Finn
Misra, Akhil
Pauli, Yves
Hüppi, Roya
Yang, Ni
Lang, Nils
Michels, Lars
Edkins, Victoria
Frühholz, Sascha
Sommer, Iris
Hinzen, Wolfram
Homan, Philipp
Neurons and Cognition
Computation and Language
To maintain coherence in language, the brain must satisfy key competing temporal demands: the gradual accumulation of meaning across extended context (drift) and the rapid reconfiguration of representations at event boundaries (shift). How these processes are implemented in the human brain during naturalistic listening remains unclear. Here, we tested whether both can be captured by annotation-free drift and shift signals and whether their neural expression shows distinct regional preferences across the brain. These signals were derived from a large language model (LLM) processing the narrative input. To enable high-precision voxelwise encoding models with stable parameter estimates, we densely sampled one healthy adult across more than 7 hours of listening to crime stories while collecting 7 Tesla fMRI data. We then modeled the feature-informed hemodynamic response using a regularized encoding framework validated on independent stories. Drift predictions were prevalent in default-mode network hubs, whereas shift predictions were evident bilaterally in the primary auditory cortex and language association cortex. Together, these findings show that coherence during language comprehension is implemented through distinct but co-expressed neural regimes of slow contextual integration and rapid event-driven reconfiguration, offering a mechanistic entry point for understanding disturbances of language coherence in psychiatric disorders.
title Coherence in the brain unfolds across separable temporal regimes
topic Neurons and Cognition
Computation and Language
url https://arxiv.org/abs/2512.20481