Instant cost and delayed reward. Demographic eco-evolutionary game dynamics under the impact of the delay resulting from the offspring maturation time

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Hauptverfasser: Argasiński, Krzysztof, Rudnicki, Ryszard, Szczelina, Robert
Format: Preprint
Veröffentlicht: 2026
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author Argasiński, Krzysztof
Rudnicki, Ryszard
Szczelina, Robert
author_facet Argasiński, Krzysztof
Rudnicki, Ryszard
Szczelina, Robert
contents In this paper, we extend the demographic eco-evolutionary game approach, based on explicit birth and death dynamics instead of abstract "fitness" interpreted as an abstract "Malthusian parameter", by the introduction of the delay resulting from the juvenile maturation time. This leads to the application of the Delay Differential Equations (DDE). We show that delay seriously affects the resulting dynamics and may lead to the loss of stability of equilibria when critical delay is exceeded. We provide theoretical tools for the assessment of the critical delays and the parameter values when this may happen. Our results emphasize the importance of the mechanisms of density dependence. We analyze the impact of three different suppression modes based on: adult mortality, juvenile recruitment survival after the maturation period (without delay), and juvenile recruitment at birth (with the delay). The last mode leads to extreme patterns such as bifurcations, complex cycles, and chaotic dynamics. However, surprisingly, this mode leads to extension of the duration of the temporary transient metastable states known as "ghost attractors". In addition, we also focus on the problem of resilience of the analyzed systems against external periodic perturbations and feedback-driven factors such as additional predator pressure.
format Preprint
id arxiv_https___arxiv_org_abs_2601_02552
institution arXiv
publishDate 2026
record_format arxiv
spellingShingle Instant cost and delayed reward. Demographic eco-evolutionary game dynamics under the impact of the delay resulting from the offspring maturation time
Argasiński, Krzysztof
Rudnicki, Ryszard
Szczelina, Robert
Populations and Evolution
Chaotic Dynamics
In this paper, we extend the demographic eco-evolutionary game approach, based on explicit birth and death dynamics instead of abstract "fitness" interpreted as an abstract "Malthusian parameter", by the introduction of the delay resulting from the juvenile maturation time. This leads to the application of the Delay Differential Equations (DDE). We show that delay seriously affects the resulting dynamics and may lead to the loss of stability of equilibria when critical delay is exceeded. We provide theoretical tools for the assessment of the critical delays and the parameter values when this may happen. Our results emphasize the importance of the mechanisms of density dependence. We analyze the impact of three different suppression modes based on: adult mortality, juvenile recruitment survival after the maturation period (without delay), and juvenile recruitment at birth (with the delay). The last mode leads to extreme patterns such as bifurcations, complex cycles, and chaotic dynamics. However, surprisingly, this mode leads to extension of the duration of the temporary transient metastable states known as "ghost attractors". In addition, we also focus on the problem of resilience of the analyzed systems against external periodic perturbations and feedback-driven factors such as additional predator pressure.
title Instant cost and delayed reward. Demographic eco-evolutionary game dynamics under the impact of the delay resulting from the offspring maturation time
topic Populations and Evolution
Chaotic Dynamics
url https://arxiv.org/abs/2601.02552