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Hauptverfasser: Zhang, Mingxin, Dai, Xiaofeng, Yao, Yu, Yin, Ziqi
Format: Preprint
Veröffentlicht: 2026
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Online-Zugang:https://arxiv.org/abs/2603.07472
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author Zhang, Mingxin
Dai, Xiaofeng
Yao, Yu
Yin, Ziqi
author_facet Zhang, Mingxin
Dai, Xiaofeng
Yao, Yu
Yin, Ziqi
contents In this study, we present a conditional diffusion-transformer framework for generating ensembles of three-dimensional Escherichia coli genome conformations guided by Hi-C contact maps. Instead of producing a single deterministic structure, we formulate genome reconstruction as a conditional generative modeling problem that samples heterogeneous conformations whose ensemble-averaged contacts are consistent with the input Hi-C data. A synthetic dataset is constructed using coarse-grained molecular dynamics simulations to generate chromatin ensembles and corresponding Hi-C maps under circular topology. Our models operate in a latent diffusion setting with a variational autoencoder that preserves per-bin alignment and supports replication-aware representations. Hi-C information is injected through a transformer-based encoder and cross-attention, enforcing a physically interpretable one-way constraint from Hi-C to structure. The model is trained using a flow-matching objective for stable optimization. On held-out ensembles, generated structures reproduce the input Hi-C distance-decay and structural correlation metrics while maintaining substantial conformational diversity, demonstrating the effectiveness of diffusion-based generative modeling for ensemble-level 3D genome reconstruction.
format Preprint
id arxiv_https___arxiv_org_abs_2603_07472
institution arXiv
publishDate 2026
record_format arxiv
spellingShingle Contact-Guided 3D Genome Structure Generation of E. coli via Diffusion Transformers
Zhang, Mingxin
Dai, Xiaofeng
Yao, Yu
Yin, Ziqi
Machine Learning
Artificial Intelligence
In this study, we present a conditional diffusion-transformer framework for generating ensembles of three-dimensional Escherichia coli genome conformations guided by Hi-C contact maps. Instead of producing a single deterministic structure, we formulate genome reconstruction as a conditional generative modeling problem that samples heterogeneous conformations whose ensemble-averaged contacts are consistent with the input Hi-C data. A synthetic dataset is constructed using coarse-grained molecular dynamics simulations to generate chromatin ensembles and corresponding Hi-C maps under circular topology. Our models operate in a latent diffusion setting with a variational autoencoder that preserves per-bin alignment and supports replication-aware representations. Hi-C information is injected through a transformer-based encoder and cross-attention, enforcing a physically interpretable one-way constraint from Hi-C to structure. The model is trained using a flow-matching objective for stable optimization. On held-out ensembles, generated structures reproduce the input Hi-C distance-decay and structural correlation metrics while maintaining substantial conformational diversity, demonstrating the effectiveness of diffusion-based generative modeling for ensemble-level 3D genome reconstruction.
title Contact-Guided 3D Genome Structure Generation of E. coli via Diffusion Transformers
topic Machine Learning
Artificial Intelligence
url https://arxiv.org/abs/2603.07472