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Main Authors: Fernández, Pamela A, Hurd, Catriona L, Roleda, Michael Y
Format: Dataset Open Access
Language:en
Published: PANGAEA 2014
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Online Access:https://doi.org/10.1594/PANGAEA.839919
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author Fernández, Pamela A
Hurd, Catriona L
Roleda, Michael Y
author_facet Fernández, Pamela A
Hurd, Catriona L
Roleda, Michael Y
collection Datos científicos de ciencias marinas y ambientales
contents Macrocystis pyrifera is a widely distributed, highly productive, seaweed. It is known to use bicarbonate (HCO3-) from seawater in photosynthesis and the main mechanism of utilization is attributed to the external catalyzed dehydration of HCO3- by the surface-bound enzyme carbonic anhydrase (CAext). Here, we examined other putative HCO3- uptake mechanisms in M. pyrifera under pHT 9.00 (HCO3-: CO2 = 940:1) and pHT 7.65 (HCO3-: CO2 = 51:1). Rates of photosynthesis, and internal CA (CAint) and CAext activity were measured following the application of AZ which inhibits CAext, and DIDS which inhibits a different HCO3- uptake system, via an anion exchange (AE) protein. We found that the main mechanism of HCO3- uptake by M. pyrifera is via an AE protein, regardless of the HCO3-: CO2 ratio, with CAext making little contribution. Inhibiting the AE protein led to a 55%-65% decrease in photosynthetic rates. Inhibiting both the AE protein and CAext at pHT 9.00 led to 80%-100% inhibition of photosynthesis, whereas at pHT 7.65, passive CO2 diffusion supported 33% of photosynthesis. CAint was active at pHT 7.65 and 9.00, and activity was always higher than CAext, because of its role in dehydrating HCO3- to supply CO2 to RuBisCO. Interestingly, the main mechanism of HCO3- uptake in M. pyrifera was different than that in other Laminariales studied (CAext-catalyzed reaction) and we suggest that species-specific knowledge of carbon uptake mechanisms is required in order to elucidate how seaweeds might respond to future changes in HCO3-:CO2 due to ocean acidification.
format Dataset Open Access
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institution PANGAEA
language en
publishDate 2014
publisher PANGAEA
record_format pangaea
spellingShingle Bicarbonate uptake via an anion exchange protein is the main mechanism of inorganic carbon acquisition by the giant kelp Macrocystis pyrifera (Laminariales, Phaeophyceae) under variable pH
Fernández, Pamela A
Hurd, Catriona L
Roleda, Michael Y
Alkalinity, total; Aragonite saturation state; Aromoana; Benthos; Bicarbonate ion; Bottles or small containers/Aquaria (<20 L); Calcite saturation state; Calculated using seacarb after Nisumaa et al. (2010); Calculated using SWCO2 (Hunter, 2007); Carbon, inorganic, dissolved; Carbonate ion; Carbonate system computation flag; Carbon dioxide; Carbonic anhydrase activity; Carbonic anhydrase activity, standard error; Chromista; Coast and continental shelf; Coulometric titration; EXP; Experiment; Fugacity of carbon dioxide (water) at sea surface temperature (wet air); Inhibition of net photosynthesis; Inhibition of net photosynthesis, standard error; Laboratory experiment; Macroalgae; Macrocystis pyrifera; Net photosynthesis rate, oxygen; Net photosynthesis rate, oxygen, standard error; OA-ICC; Ocean acidification; Ocean Acidification International Coordination Centre; Ochrophyta; Other metabolic rates; Partial pressure of carbon dioxide (water) at sea surface temperature (wet air); pH, total scale; Potentiometric titration; Primary production/Photosynthesis; Salinity; Single species; South Pacific; Species; Spectrophotometric; Temperate; Temperature, water
Macrocystis pyrifera is a widely distributed, highly productive, seaweed. It is known to use bicarbonate (HCO3-) from seawater in photosynthesis and the main mechanism of utilization is attributed to the external catalyzed dehydration of HCO3- by the surface-bound enzyme carbonic anhydrase (CAext). Here, we examined other putative HCO3- uptake mechanisms in M. pyrifera under pHT 9.00 (HCO3-: CO2 = 940:1) and pHT 7.65 (HCO3-: CO2 = 51:1). Rates of photosynthesis, and internal CA (CAint) and CAext activity were measured following the application of AZ which inhibits CAext, and DIDS which inhibits a different HCO3- uptake system, via an anion exchange (AE) protein. We found that the main mechanism of HCO3- uptake by M. pyrifera is via an AE protein, regardless of the HCO3-: CO2 ratio, with CAext making little contribution. Inhibiting the AE protein led to a 55%-65% decrease in photosynthetic rates. Inhibiting both the AE protein and CAext at pHT 9.00 led to 80%-100% inhibition of photosynthesis, whereas at pHT 7.65, passive CO2 diffusion supported 33% of photosynthesis. CAint was active at pHT 7.65 and 9.00, and activity was always higher than CAext, because of its role in dehydrating HCO3- to supply CO2 to RuBisCO. Interestingly, the main mechanism of HCO3- uptake in M. pyrifera was different than that in other Laminariales studied (CAext-catalyzed reaction) and we suggest that species-specific knowledge of carbon uptake mechanisms is required in order to elucidate how seaweeds might respond to future changes in HCO3-:CO2 due to ocean acidification.
title Bicarbonate uptake via an anion exchange protein is the main mechanism of inorganic carbon acquisition by the giant kelp Macrocystis pyrifera (Laminariales, Phaeophyceae) under variable pH
topic Alkalinity, total; Aragonite saturation state; Aromoana; Benthos; Bicarbonate ion; Bottles or small containers/Aquaria (<20 L); Calcite saturation state; Calculated using seacarb after Nisumaa et al. (2010); Calculated using SWCO2 (Hunter, 2007); Carbon, inorganic, dissolved; Carbonate ion; Carbonate system computation flag; Carbon dioxide; Carbonic anhydrase activity; Carbonic anhydrase activity, standard error; Chromista; Coast and continental shelf; Coulometric titration; EXP; Experiment; Fugacity of carbon dioxide (water) at sea surface temperature (wet air); Inhibition of net photosynthesis; Inhibition of net photosynthesis, standard error; Laboratory experiment; Macroalgae; Macrocystis pyrifera; Net photosynthesis rate, oxygen; Net photosynthesis rate, oxygen, standard error; OA-ICC; Ocean acidification; Ocean Acidification International Coordination Centre; Ochrophyta; Other metabolic rates; Partial pressure of carbon dioxide (water) at sea surface temperature (wet air); pH, total scale; Potentiometric titration; Primary production/Photosynthesis; Salinity; Single species; South Pacific; Species; Spectrophotometric; Temperate; Temperature, water
url https://doi.org/10.1594/PANGAEA.839919