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Apopylus Kolibac 2003

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1. Verfasser:	Bartlett, Justin S.
Format:	Recurso digital
Veröffentlicht:	Zenodo 2024
Schlagworte:	Biodiversity Taxonomy Animalia Arthropoda Insecta Coleoptera Cleridae Apopylus
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author	Bartlett, Justin S.
author_facet	Bartlett, Justin S.
contents	<p><b><i>Apopylus</i> Kolibá&ccaron;, 2003</b></p><p><i>Apopylus</i> Kolibá&ccaron;, 2003: 65. Type species: <i>Apopylus unumgarensis</i> Kolibá&ccaron;, 2003: 67, by original designation. Gender: Masculine.</p><p><b>Revised differential diagnosis.</b> <i>Apopylus</i> is differentiated from <i>Parapylus</i> and <i>Pylus</i> as follows: Eyes margin flush with cranium, or only finely and unevenly margined (Figs. 1, 2), not with a clearly raised margin as in <i>Pylus</i> and <i>Parapylus</i>; antennal club loose (Figs 6–8) (compact in <i>Pylus</i> and <i>Parapylus</i>), segments 10 and 11 not abruptly cupuliform (as in <i>Pylus</i> and <i>Parapylus</i>); terminal palpomeres notably large in relation to head size (Fig. 14) (significantly smaller in <i>Pylus</i> and <i>Parapylus</i>); prothorax weakly (Fig. 16) (most species) to moderately (only <i>A. leptofustus</i> Opitz comb. nov.) tuberculate laterally (strongly tuberculate in <i>Pylus</i> and <i>Parapylus</i>), pronotal disc with matrix of shallow crater-like impressions (Fig. 16) (<i>Pylus</i> and <i>Parapylus</i> with more isolated deep foveolate punctures), disc also without central sulcus (disc sulcate in most <i>Pylus</i> and <i>Parapylus</i> species); mesocoxal cavities narrowly open to completely closed laterally (typically wide open in <i>Parapylus</i> and <i>Pylus</i>); elytral margin distally with curved flange and minute bead-like nodules (Fig. 19) (beading absent in <i>Pylus</i> and <i>Parapylus</i>), discal punctures always with four small internal nodules, i.e., quadrinodal (Fig. 20) (two in <i>Pylus</i> and <i>Parapylus</i>).</p><p>Putative synapomorphies supporting monophyly of <i>Apopylus</i> in relation to <i>Pylus</i> and <i>Parapylus</i> include the enlargement of terminal palpomeres, the narrowing (or complete closure) of the mesocoxal cavities, and the development of beading along the apical elytral margin. Whether four internal nodules of the elytral punctures represents the apomorphic or plesiomorphic state (in relation to the two nodules of <i>Pylus</i> and <i>Parapylus</i>) is less clear.</p><p><i>......Figure legend continued on the next page</i></p><p>Characters variable within <i>Apopylus</i> include: antennal club segments notably elongate (most species), more compact (<i>A. creperus</i> comb. nov.), or intermediate (<i>A. unumgarensis</i>); frons about one eye width or slightly narrower (<i>A. astrictus</i> comb. nov., <i>A. leptofustus</i> comb. nov., <i>A. redactus</i> comb. nov.) or almost two eye widths (<i>A. unumgarensis, A. creperus</i> comb. nov.); pronotal lateral tubercle very weak (most species) or moderate (<i>A. leptofustus</i> comb. nov.); mesocoxal cavities closed (<i>A. unumgarensis</i> and <i>A. creperus</i> comb. nov.) or very narrowly open (<i>A. leptofustus</i> comb. nov.) laterally (not documented in <i>A. astrictus</i> comb. nov. or <i>A. redactus</i> comb. nov.); tibial spur formulae 1–2–1 (most species) or 0–0–0 (<i>A. unumgarensis</i>).</p><p><b>Included species.</b> (5): <i>Apopylus unumgarensis</i> Kolibá&ccaron;,2003; <i>Apopylus astrictus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>); <i>Apopylus creperus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>); <i>Apopylus leptofustus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>); <i>Apopylus redactus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>).</p><p><b>Material examined.</b> <i>Apopylus unumgarensis</i> Kolibá&ccaron; (4): NSW, Unumgar S.F., 430 m, nr Woodenbong, Pole Bridge Road, 788, 2–11 Jan 1987, A. Newton & M. Thayer (holotype, ANIC); QLD: 26.881&ring;Sx151.600&ring;E Bunyas, Dandabah, 1000m, 2Oct 2010, G. Monteith, RF barkspray, 34628 (2, JSBC); SEQ:24°32’Sx151°28’E Bulburin barracks. 8 Oct 1999. 580 m. G.B.Monteith. rainforest Pyrtethrum, trees. 7816 // <i>Apopylus unumgarensis</i> Kolibá&ccaron; Det. W. Opitz (1, QM).</p><p><i>Fallopylus creperus</i> Opitz (2): Australia: n. Qld. 7km NE of Tolga. Nov 1987, Storey & Defaveri. Light trap (1 paratype, QDPC); same data as above but 10.xii.1986 // Fallopylus creperus Det. JS Bartlett 2017 (1, QDPC).</p><p><i>Fallopylus leptofustus</i> Opitz (3): Australia: n. Qld. Tully Falls S.F. 730 m. 18 km SSW Ravenshoe 1.x. – 5.xi.1987. Storey & Dickenson (1 &female; paratype, QDPC); Australia: n. Qld. Wongabel S.F. via Atherton. 4-16.i.1990. Story & Defaveri. Malaise trap (1 paratype, QDPC); NEQ:17°33’Sx145°33’E Mt Fisher, summit. 1360m 8 Feb 1999. Rainforest GB Monteith. Pyrethrum —trees & logs 2176 (1 paratype, QM).</p><p><i>Fallopylus redactus</i> Opitz (1): Tamborine Mt., Joalah NP, NSW, 25 Nov. 1982, J. & E. Doyen (holotype, female, ANIC).</p><p><b>Remarks.</b> Specimens of <i>Fallopylus astrictus</i> Opitz were not examined. It’s transfer to <i>Apopylus</i> is based on Opitz’ (2015) description of the antennal capitulum as loose, the pronotal disc as not centrally impressed, the elytral epipleurae as distally serrated, the elytral punctation as quadrinodal and the pronotal side margins gradually broadening and not prominently tuberculate (Opitz 2015, Fig. 67).</p><p>Opitz (2015) described the elytral punctures of <i>Apopylus unumgarensis</i> as binodal and those of <i>A. creparus</i> as trinodal; examination of type specimens revealed that the punctures are quadrinodal in both species.</p><p>To this author’s knowledge, the phenomenon of inconsistent expression of number of apical tibial spurs among the species of a clerid genus is known only in the genera <i>Apopylus</i> and <i>Pylus.</i></p>
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spellingShingle	Apopylus Kolibac 2003 Bartlett, Justin S. Biodiversity Taxonomy Animalia Arthropoda Insecta Coleoptera Cleridae Apopylus <p><b><i>Apopylus</i> Kolibá&ccaron;, 2003</b></p><p><i>Apopylus</i> Kolibá&ccaron;, 2003: 65. Type species: <i>Apopylus unumgarensis</i> Kolibá&ccaron;, 2003: 67, by original designation. Gender: Masculine.</p><p><b>Revised differential diagnosis.</b> <i>Apopylus</i> is differentiated from <i>Parapylus</i> and <i>Pylus</i> as follows: Eyes margin flush with cranium, or only finely and unevenly margined (Figs. 1, 2), not with a clearly raised margin as in <i>Pylus</i> and <i>Parapylus</i>; antennal club loose (Figs 6–8) (compact in <i>Pylus</i> and <i>Parapylus</i>), segments 10 and 11 not abruptly cupuliform (as in <i>Pylus</i> and <i>Parapylus</i>); terminal palpomeres notably large in relation to head size (Fig. 14) (significantly smaller in <i>Pylus</i> and <i>Parapylus</i>); prothorax weakly (Fig. 16) (most species) to moderately (only <i>A. leptofustus</i> Opitz comb. nov.) tuberculate laterally (strongly tuberculate in <i>Pylus</i> and <i>Parapylus</i>), pronotal disc with matrix of shallow crater-like impressions (Fig. 16) (<i>Pylus</i> and <i>Parapylus</i> with more isolated deep foveolate punctures), disc also without central sulcus (disc sulcate in most <i>Pylus</i> and <i>Parapylus</i> species); mesocoxal cavities narrowly open to completely closed laterally (typically wide open in <i>Parapylus</i> and <i>Pylus</i>); elytral margin distally with curved flange and minute bead-like nodules (Fig. 19) (beading absent in <i>Pylus</i> and <i>Parapylus</i>), discal punctures always with four small internal nodules, i.e., quadrinodal (Fig. 20) (two in <i>Pylus</i> and <i>Parapylus</i>).</p><p>Putative synapomorphies supporting monophyly of <i>Apopylus</i> in relation to <i>Pylus</i> and <i>Parapylus</i> include the enlargement of terminal palpomeres, the narrowing (or complete closure) of the mesocoxal cavities, and the development of beading along the apical elytral margin. Whether four internal nodules of the elytral punctures represents the apomorphic or plesiomorphic state (in relation to the two nodules of <i>Pylus</i> and <i>Parapylus</i>) is less clear.</p><p><i>......Figure legend continued on the next page</i></p><p>Characters variable within <i>Apopylus</i> include: antennal club segments notably elongate (most species), more compact (<i>A. creperus</i> comb. nov.), or intermediate (<i>A. unumgarensis</i>); frons about one eye width or slightly narrower (<i>A. astrictus</i> comb. nov., <i>A. leptofustus</i> comb. nov., <i>A. redactus</i> comb. nov.) or almost two eye widths (<i>A. unumgarensis, A. creperus</i> comb. nov.); pronotal lateral tubercle very weak (most species) or moderate (<i>A. leptofustus</i> comb. nov.); mesocoxal cavities closed (<i>A. unumgarensis</i> and <i>A. creperus</i> comb. nov.) or very narrowly open (<i>A. leptofustus</i> comb. nov.) laterally (not documented in <i>A. astrictus</i> comb. nov. or <i>A. redactus</i> comb. nov.); tibial spur formulae 1–2–1 (most species) or 0–0–0 (<i>A. unumgarensis</i>).</p><p><b>Included species.</b> (5): <i>Apopylus unumgarensis</i> Kolibá&ccaron;,2003; <i>Apopylus astrictus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>); <i>Apopylus creperus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>); <i>Apopylus leptofustus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>); <i>Apopylus redactus</i> (Opitz, 2015) <b>new combination</b> (transferred from <i>Fallopylus</i>).</p><p><b>Material examined.</b> <i>Apopylus unumgarensis</i> Kolibá&ccaron; (4): NSW, Unumgar S.F., 430 m, nr Woodenbong, Pole Bridge Road, 788, 2–11 Jan 1987, A. Newton & M. Thayer (holotype, ANIC); QLD: 26.881&ring;Sx151.600&ring;E Bunyas, Dandabah, 1000m, 2Oct 2010, G. Monteith, RF barkspray, 34628 (2, JSBC); SEQ:24°32’Sx151°28’E Bulburin barracks. 8 Oct 1999. 580 m. G.B.Monteith. rainforest Pyrtethrum, trees. 7816 // <i>Apopylus unumgarensis</i> Kolibá&ccaron; Det. W. Opitz (1, QM).</p><p><i>Fallopylus creperus</i> Opitz (2): Australia: n. Qld. 7km NE of Tolga. Nov 1987, Storey & Defaveri. Light trap (1 paratype, QDPC); same data as above but 10.xii.1986 // Fallopylus creperus Det. JS Bartlett 2017 (1, QDPC).</p><p><i>Fallopylus leptofustus</i> Opitz (3): Australia: n. Qld. Tully Falls S.F. 730 m. 18 km SSW Ravenshoe 1.x. – 5.xi.1987. Storey & Dickenson (1 &female; paratype, QDPC); Australia: n. Qld. Wongabel S.F. via Atherton. 4-16.i.1990. Story & Defaveri. Malaise trap (1 paratype, QDPC); NEQ:17°33’Sx145°33’E Mt Fisher, summit. 1360m 8 Feb 1999. Rainforest GB Monteith. Pyrethrum —trees & logs 2176 (1 paratype, QM).</p><p><i>Fallopylus redactus</i> Opitz (1): Tamborine Mt., Joalah NP, NSW, 25 Nov. 1982, J. & E. Doyen (holotype, female, ANIC).</p><p><b>Remarks.</b> Specimens of <i>Fallopylus astrictus</i> Opitz were not examined. It’s transfer to <i>Apopylus</i> is based on Opitz’ (2015) description of the antennal capitulum as loose, the pronotal disc as not centrally impressed, the elytral epipleurae as distally serrated, the elytral punctation as quadrinodal and the pronotal side margins gradually broadening and not prominently tuberculate (Opitz 2015, Fig. 67).</p><p>Opitz (2015) described the elytral punctures of <i>Apopylus unumgarensis</i> as binodal and those of <i>A. creparus</i> as trinodal; examination of type specimens revealed that the punctures are quadrinodal in both species.</p><p>To this author’s knowledge, the phenomenon of inconsistent expression of number of apical tibial spurs among the species of a clerid genus is known only in the genera <i>Apopylus</i> and <i>Pylus.</i></p>
title	Apopylus Kolibac 2003
topic	Biodiversity Taxonomy Animalia Arthropoda Insecta Coleoptera Cleridae Apopylus
url	https://doi.org/10.5281/zenodo.10708563