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Table 3 in Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species

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Hauptverfasser:	Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo, Gallao 2, Jonas E.
Format:	Recurso digital
Veröffentlicht:	Zenodo 2021
Schlagworte:	Biodiversity Taxonomy
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_version_	1866902014204575744
author	Moreno-Gonzalez 1, Jairo A. Pinto-da-Rocha 1, Ricardo Gallao 2, Jonas E.
author_facet	Moreno-Gonzalez 1, Jairo A. Pinto-da-Rocha 1, Ricardo Gallao 2, Jonas E.
contents	<p><b>Table 3.</b> Phenotypic characters useful for the taxonomy of <i>Tityus</i>. (*) Species here transferred to the indicated subgenus; ventral macrosetae distribution on telotarsi I-IV: <b>Type I</b> = tuft of irregularly distributed macrosetae. <b>Type II</b> = two discrete ventrosubmedian rows of macrosetae. Abbreviations: BML, basal middle lamellae; D, dilated; NA, not applicable; ND, not dilated; PBP, pectinal basal piece.</p><table><thead><tr><th colspan="1" rowspan="2">Species</th><th colspan="1" rowspan="2">Subgenus</th><th colspan="1" rowspan="2">Species Group</th><th colspan="1" rowspan="2">Telotarsal setae</th><th colspan="2" rowspan="1">Females</th></tr><tr><th colspan="1" rowspan="1">PBP gland</th><th colspan="1" rowspan="1">BML</th></tr></thead><tbody><tr><td colspan="1" rowspan="1"><i>Isometrus maculatus</i> (DeGeer, 1778)</td><td colspan="1" rowspan="1">NA</td><td colspan="1" rowspan="1">NA</td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus argentinus</i> Borelli, 1899</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus bahiensis</i> (Perty, 1833)</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus blaseri</i> Mello-Leitão, 1931</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">More than first 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus brazilae</i> Lourenço & Eickstedt, 1984</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus carrilloi</i> Ojanguren-Affilastro, 2021</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus carvalhoi</i> Mello-Leitão, 1945</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus charreyroni</i> Mello-Leitão, 1933</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus clathratus</i> C. L. Koch, 1844</td><td colspan="1" rowspan="1"><i>Archaeotityus</i></td><td colspan="1" rowspan="1"><i>T. clathratus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">First anteromedian third</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus confluens</i> Borelli, 1899</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus curupi</i> Ojanguren-Affilastro et al. 2017</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= suboval</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus forcipula</i> (Gervais, 1843)</td><td colspan="1" rowspan="1"><i>Atreus</i></td><td colspan="1" rowspan="1"><i>T. forcipula</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= suboval</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus obscurus</i> Gervais, 1843</td><td colspan="1" rowspan="1"><i>Atreus</i></td><td colspan="1" rowspan="1"><i>T. obscurus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus panguana</i> Kova&rcaron;ík et al. 2015</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus potameis</i> Lourenço & Giupponi, 2004</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus sastrei</i> Lourenço & Flórez, 1990**</td><td colspan="1" rowspan="1"><i>Atreus</i></td><td colspan="1" rowspan="1">NA</td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus serrulatus</i> Lutz & Melo, 1922</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. stigmurus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First anteromedian third</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus soratensis</i> Kraepelin, 1912</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">?</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus spelaeus</i> sp. nov.</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">More than first 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus stigmurus</i> (Thorell, 1876)</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. stigmurus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First anteromedian third</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus trivittatus</i> Kraepelin, 1898</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus uruguayensis</i> Borelli, 1901</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr></tbody></table>
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id	zenodo_https___doi_org_10_5281_zenodo_15143501
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publishDate	2021
publisher	Zenodo
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spellingShingle	Table 3 in Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species Moreno-Gonzalez 1, Jairo A. Pinto-da-Rocha 1, Ricardo Gallao 2, Jonas E. Biodiversity Taxonomy <p><b>Table 3.</b> Phenotypic characters useful for the taxonomy of <i>Tityus</i>. (*) Species here transferred to the indicated subgenus; ventral macrosetae distribution on telotarsi I-IV: <b>Type I</b> = tuft of irregularly distributed macrosetae. <b>Type II</b> = two discrete ventrosubmedian rows of macrosetae. Abbreviations: BML, basal middle lamellae; D, dilated; NA, not applicable; ND, not dilated; PBP, pectinal basal piece.</p><table><thead><tr><th colspan="1" rowspan="2">Species</th><th colspan="1" rowspan="2">Subgenus</th><th colspan="1" rowspan="2">Species Group</th><th colspan="1" rowspan="2">Telotarsal setae</th><th colspan="2" rowspan="1">Females</th></tr><tr><th colspan="1" rowspan="1">PBP gland</th><th colspan="1" rowspan="1">BML</th></tr></thead><tbody><tr><td colspan="1" rowspan="1"><i>Isometrus maculatus</i> (DeGeer, 1778)</td><td colspan="1" rowspan="1">NA</td><td colspan="1" rowspan="1">NA</td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus argentinus</i> Borelli, 1899</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus bahiensis</i> (Perty, 1833)</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus blaseri</i> Mello-Leitão, 1931</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">More than first 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus brazilae</i> Lourenço & Eickstedt, 1984</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus carrilloi</i> Ojanguren-Affilastro, 2021</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus carvalhoi</i> Mello-Leitão, 1945</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus charreyroni</i> Mello-Leitão, 1933</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bahiensis</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus clathratus</i> C. L. Koch, 1844</td><td colspan="1" rowspan="1"><i>Archaeotityus</i></td><td colspan="1" rowspan="1"><i>T. clathratus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">First anteromedian third</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus confluens</i> Borelli, 1899</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus curupi</i> Ojanguren-Affilastro et al. 2017</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= suboval</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus forcipula</i> (Gervais, 1843)</td><td colspan="1" rowspan="1"><i>Atreus</i></td><td colspan="1" rowspan="1"><i>T. forcipula</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= suboval</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus obscurus</i> Gervais, 1843</td><td colspan="1" rowspan="1"><i>Atreus</i></td><td colspan="1" rowspan="1"><i>T. obscurus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus panguana</i> Kova&rcaron;ík et al. 2015</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus potameis</i> Lourenço & Giupponi, 2004</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus sastrei</i> Lourenço & Flórez, 1990**</td><td colspan="1" rowspan="1"><i>Atreus</i></td><td colspan="1" rowspan="1">NA</td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus serrulatus</i> Lutz & Melo, 1922</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. stigmurus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First anteromedian third</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus soratensis</i> Kraepelin, 1912</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">?</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus spelaeus</i> sp. nov.</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">More than first 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus stigmurus</i> (Thorell, 1876)</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. stigmurus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First anteromedian third</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus trivittatus</i> Kraepelin, 1898</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. trivittatus</i></td><td colspan="1" rowspan="1">Type II</td><td colspan="1" rowspan="1">First 2/3 of the anterior region</td><td colspan="1" rowspan="1">ND</td></tr><tr><td colspan="1" rowspan="1"><i>Tityus uruguayensis</i> Borelli, 1901</td><td colspan="1" rowspan="1"><i>Tityus</i></td><td colspan="1" rowspan="1"><i>T. bolivianus</i></td><td colspan="1" rowspan="1">Type I</td><td colspan="1" rowspan="1">Absent</td><td colspan="1" rowspan="1">D= semicircular</td></tr></tbody></table>
title	Table 3 in Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species
topic	Biodiversity Taxonomy
url	https://doi.org/10.5281/zenodo.15143501