Dichotomius (Dichotomius) boreus

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Hauptverfasser: Arias-Buriticá, Jorge Armando, Vaz-de-Mello, Fernando Zagury
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Veröffentlicht: Zenodo 2025
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author Arias-Buriticá, Jorge Armando
Vaz-de-Mello, Fernando Zagury
author_facet Arias-Buriticá, Jorge Armando
Vaz-de-Mello, Fernando Zagury
contents <p><b>2. Dichotomius (Dichotomius) boreus (Olivier, 1789) species group</b></p><p><b>Diagnosis:</b> The species of the D. boreus species group are recognized by the following combination of characters: (1) Medium-sized to large-sized individuals (16.6–27.1 mm). (2) Head oval and twice as wide as it is long (Fig. 7a, b). (3) Clypeus with two inconspicuous clypeal teeth, sometimes imperceptible (Fig. 7a, b). (4) Ventral clypeal process with invagination and then a conical process obtuse or acute tip (Fig. 7c). (5) Both sexes with two cephalic process, the first one located in the clypeal region and the second one in the fronto-clypeal region (Fig. 7a, b). (6) Protibial spur not angled or slightly angled near the apex (Fig. 7d). (7) Pronotum with an excavation in the anterior area that can go from 1/3 to half of it and laterally to more than 2/3 of the pronotum (Fig. 7a, b). (8) Pronotum with central process with different shapes according to the species. (9) Pronotal process with two humps with spine shape above the lateral foveae. (10) Posterior margin of pronotum with ocellate punctures, sometimes denser in the posteromedial area (Figs.2a, b, 7a, b). (11) Elytral striae bicarinate, shallow to distinctly impressed and with punctate interestriae. (12) Pygidium with complete apical margin and evident punctures (Fig. 7e) (13). Aedeagus lacking subgenital plate (Fig. 7h). (14) Parameres subtriangular in lateral view (Fig. 7f), without setae in the apical area (Fig. 7 f-h). (15) ME well sclerotized, large, and asymmetrical (Fig. 7l).</p><p><b>Composition:</b> Six valid species: Dichotomius boreus (Olivier, 1789), D. carinatus (Luederwaldt, 1925), D.podalirius (Felsche, 1901), D. prietoi Martínez & Martínez, 1982, D. pelamon (Harold, 1869), D.quadrilobatus Chamorro et al., 2021, and two undescribed species.</p><p><b>Geographic distribution:</b> Pacific dominion (Sabana province), Boreal brazilian dominion (all provinces), South Brazilian dominion (all provinces), Chacoan subregion Southeastern Amazonian dominion (Xingu-Tapajós province), and South American Transition zones (Páramo province) (Fig. 6b).</p>
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id zenodo_https___doi_org_10_5281_zenodo_16975099
institution Zenodo
language
publishDate 2025
publisher Zenodo
record_format zenodo
spellingShingle Dichotomius (Dichotomius) boreus
Arias-Buriticá, Jorge Armando
Vaz-de-Mello, Fernando Zagury
Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Dichotomius
Dichotomius boreus
<p><b>2. Dichotomius (Dichotomius) boreus (Olivier, 1789) species group</b></p><p><b>Diagnosis:</b> The species of the D. boreus species group are recognized by the following combination of characters: (1) Medium-sized to large-sized individuals (16.6–27.1 mm). (2) Head oval and twice as wide as it is long (Fig. 7a, b). (3) Clypeus with two inconspicuous clypeal teeth, sometimes imperceptible (Fig. 7a, b). (4) Ventral clypeal process with invagination and then a conical process obtuse or acute tip (Fig. 7c). (5) Both sexes with two cephalic process, the first one located in the clypeal region and the second one in the fronto-clypeal region (Fig. 7a, b). (6) Protibial spur not angled or slightly angled near the apex (Fig. 7d). (7) Pronotum with an excavation in the anterior area that can go from 1/3 to half of it and laterally to more than 2/3 of the pronotum (Fig. 7a, b). (8) Pronotum with central process with different shapes according to the species. (9) Pronotal process with two humps with spine shape above the lateral foveae. (10) Posterior margin of pronotum with ocellate punctures, sometimes denser in the posteromedial area (Figs.2a, b, 7a, b). (11) Elytral striae bicarinate, shallow to distinctly impressed and with punctate interestriae. (12) Pygidium with complete apical margin and evident punctures (Fig. 7e) (13). Aedeagus lacking subgenital plate (Fig. 7h). (14) Parameres subtriangular in lateral view (Fig. 7f), without setae in the apical area (Fig. 7 f-h). (15) ME well sclerotized, large, and asymmetrical (Fig. 7l).</p><p><b>Composition:</b> Six valid species: Dichotomius boreus (Olivier, 1789), D. carinatus (Luederwaldt, 1925), D.podalirius (Felsche, 1901), D. prietoi Martínez & Martínez, 1982, D. pelamon (Harold, 1869), D.quadrilobatus Chamorro et al., 2021, and two undescribed species.</p><p><b>Geographic distribution:</b> Pacific dominion (Sabana province), Boreal brazilian dominion (all provinces), South Brazilian dominion (all provinces), Chacoan subregion Southeastern Amazonian dominion (Xingu-Tapajós province), and South American Transition zones (Páramo province) (Fig. 6b).</p>
title Dichotomius (Dichotomius) boreus
topic Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Dichotomius
Dichotomius boreus
url https://doi.org/10.5281/zenodo.16975099