Paraphytini Scholtz, Davis & de Klerk 2025, new tribe
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| author | Scholtz, Clarke H. Davis, Adrian L. V. Klerk, Hennie De |
| author_facet | Scholtz, Clarke H. Davis, Adrian L. V. Klerk, Hennie De |
| contents | <p><b>Paraphytini Scholtz, Davis & de Klerk, new tribe</b></p><p><b>Type genus</b>: <i>Paraphytus</i> Harold, 1877, here designated (Fig. 6)</p><p><b>Diagnosis</b></p><p>Differs from other tribes by a combination of characters, primarily a habitus similar to that of the sister subfamily, Aphodiinae combined with a quadridentate clypeus and apical excavation of the tridentate protibiae.</p><p><b>Description</b></p><p><b>Head:</b> Anterior clypeal margin quadridentate.</p><p><b>Thorax:</b> Pronotal disc punctate and evenly convex lacking carinae or other protrusions.</p><p><b>Wings:</b> Vein characters of metathoracic wing not modified (Fig. 7a) compared to those shown in Fig. 7b: (1) anal area not reduced, (2) no widening of CuA vein at wing margin, (3) RA 4 vein reaches wing margin. Elytra punctate and evenly convex with seven dorsal striae and an eighth short stria positioned laterally. Epipleurae only very weakly sinuate (see Fig. 34h in Tarasov & Génier 2015).</p><p><b>Legs:</b> Short tarsi inserted on the upper side of the tridentate fore tibiae (Tarasov & Génier 2015) within an apical excavation.</p><p><b>Abdomen:</b> Pygidium longitudinally grooved in the centre or not (see Paulian 1936).</p><p><b>Male genitalia:</b> Using the terminology of Tarasov & Solodovnikov (2011) and Tarasov & Génier (2015) for the complex of accessory sclerites occurring within the basal part of aedeagal sacs (in general: A, AS, BSc, FLP, SA, SRP and X), <i>Paraphytus</i> shows: (1) a long spur (=TS) and a short lobe extending from the A sclerite, (2) presence of a BSc sclerite and (3) absence of other sclerites (Tarasov & Génier 2015: p. 36). However, somewhat similar characterization of the endophallites is also shared with genera of the Coptorhinini <b>new tribe</b> (see below) and some other “basal Scarabaeinae ” (Tarasov & Dimitrov 2016) that are now in the tribes Elassocanthonini or Endroedyolini (Tarasov & Génier 2015: pp. 35–37).</p><p><b>Sexual dimorphism:</b> Limited to differences in the sculpture of the pygidium, abdominal sternites and clypeal margin.</p> |
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| id | zenodo_https___doi_org_10_5281_zenodo_17884243 |
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| publishDate | 2025 |
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| spellingShingle | Paraphytini Scholtz, Davis & de Klerk 2025, new tribe Scholtz, Clarke H. Davis, Adrian L. V. Klerk, Hennie De Biodiversity Taxonomy Animalia Arthropoda Insecta Coleoptera Scarabaeidae <p><b>Paraphytini Scholtz, Davis & de Klerk, new tribe</b></p><p><b>Type genus</b>: <i>Paraphytus</i> Harold, 1877, here designated (Fig. 6)</p><p><b>Diagnosis</b></p><p>Differs from other tribes by a combination of characters, primarily a habitus similar to that of the sister subfamily, Aphodiinae combined with a quadridentate clypeus and apical excavation of the tridentate protibiae.</p><p><b>Description</b></p><p><b>Head:</b> Anterior clypeal margin quadridentate.</p><p><b>Thorax:</b> Pronotal disc punctate and evenly convex lacking carinae or other protrusions.</p><p><b>Wings:</b> Vein characters of metathoracic wing not modified (Fig. 7a) compared to those shown in Fig. 7b: (1) anal area not reduced, (2) no widening of CuA vein at wing margin, (3) RA 4 vein reaches wing margin. Elytra punctate and evenly convex with seven dorsal striae and an eighth short stria positioned laterally. Epipleurae only very weakly sinuate (see Fig. 34h in Tarasov & Génier 2015).</p><p><b>Legs:</b> Short tarsi inserted on the upper side of the tridentate fore tibiae (Tarasov & Génier 2015) within an apical excavation.</p><p><b>Abdomen:</b> Pygidium longitudinally grooved in the centre or not (see Paulian 1936).</p><p><b>Male genitalia:</b> Using the terminology of Tarasov & Solodovnikov (2011) and Tarasov & Génier (2015) for the complex of accessory sclerites occurring within the basal part of aedeagal sacs (in general: A, AS, BSc, FLP, SA, SRP and X), <i>Paraphytus</i> shows: (1) a long spur (=TS) and a short lobe extending from the A sclerite, (2) presence of a BSc sclerite and (3) absence of other sclerites (Tarasov & Génier 2015: p. 36). However, somewhat similar characterization of the endophallites is also shared with genera of the Coptorhinini <b>new tribe</b> (see below) and some other “basal Scarabaeinae ” (Tarasov & Dimitrov 2016) that are now in the tribes Elassocanthonini or Endroedyolini (Tarasov & Génier 2015: pp. 35–37).</p><p><b>Sexual dimorphism:</b> Limited to differences in the sculpture of the pygidium, abdominal sternites and clypeal margin.</p> |
| title | Paraphytini Scholtz, Davis & de Klerk 2025, new tribe |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Coleoptera Scarabaeidae |
| url | https://doi.org/10.5281/zenodo.17884243 |