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2025
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| author | Alzahrani, Ali Mohammed Magos Brehm, Joana Ghazanfar, Shahina A. Maxted, Nigel |
| author_facet | Alzahrani, Ali Mohammed Magos Brehm, Joana Ghazanfar, Shahina A. Maxted, Nigel |
| contents | <p><b>15.</b></p><i>Verbascum transjordanicum</i>Murb. in Lunds Univ. Arsskrift, n. f. xxxv. No. 1 54 (1939)<p>Fig. 28</p><p><b>Type.</b></p><p>Jordan, Sandy desert, 40 km west of Azrak, 17 April 1936, <i>J. E. Dinsmore 11804</i> (isotype E [E 00327349]).</p><p><b>Description.</b></p><p>Annual or biennial herb, pale green, usually branched, many-stemmed from the base, up to 70 cm tall. <i>Indumentum</i> dense glandular hairs with sparse simple and forked hairs above, and dense tomentose with stellate hairs below. <i>Stems</i> erect, robust, terete to angular. <i>Basal leaves</i> rosette, oblong-lanceolate, 2.5–12 × 1–3.5 cm, apex acute, base obtuse, margins crenate to pinnatifid-lobed, lamina darkish green with sparse stellate hairs above and dense white-grey tomentose with stellate hairs below; petiole 1–4 cm. <i>Cauline leaves</i> oblong-lanceolate, 1.6–5 × 1–2 cm, apex acute, base obtuse; sessile. <i>Inflorescence</i> racemose; flowers single in the axil of bracts. <i>Upper bracts</i> ovate-triangular, 1–4 mm, acute. <i>Lower bracts</i> oblong-lanceolate, 4–8 mm, acute. <i>Pedicel</i> covered with dense glandular hairs up to 20 mm long. <i>Bracteoles</i> absent. <i>Calyx</i> 3.4–4.4 mm, lobes oblong-lanceolate, acute, dense glandular. <i>Corolla</i> 20–30 mm across, yellow with dark purple-red hairy blotches around the throat, without pellucid glands, tube up to 1 mm, dense papillose hairs inside, spare glandular, simple, and forked hairs outside. <i>Stamens</i> five, 3–5 mm long. <i>Filaments</i> yellow with creamy hairs, two anterior glabrous near the apex, three posteriors with hairs up to anthers. <i>Anthers</i> all reniform. <i>Ovary</i> globose-ovoid, sparse glandular hairs. <i>Style</i> up to 7 mm long, filiform, green. <i>Stigma</i> capitate. <i>Capsule</i> 4–6 × 3–4 mm, globose-ovoid, sparse glandular hairs. <i>Seeds</i> bothrospermous.</p><p><b>Distribution.</b></p><p>Jordan and Saudi Arabia.</p><p><b>Distribution in the Arabian Peninsula.</b></p><p>It is a native species to Saudi Arabia, which is known from Turaif province (near Harrat Al-Harrat Reserve), northern Saudi Arabia (Fig. 29).</p><p><b>Ecology.</b></p><p><i>Verbascum transjordanicum</i> grows on a limestone plateau with basalt rock at altitudes ranging from 600 to 832 m. Associated plants include <i>Achillea fragrantissima</i> (Forssk.) Sch. Bip., <i>Centaurea sinaica</i> DC., <i>Cornulaca setifera</i> (DC.) Moq., <i>Helianthemum lippii</i> (L.) Dum. Cours., and <i>Hyoscyamus muticus subsp. muticus</i>.</p><p><b>Vernacular name.</b></p><p><i>Transjordan Mullein</i> (English), <i>Abu Ain</i> (Arabic).</p><p><b>Phenology.</b></p><p>Flowering and fruiting from March to June.</p><p><b>Etymology.</b></p><p>The name refers to its occurrence in the Transjordan region, meaning “ across ” or “ beyond ” Jordan.</p><p><b>Specimens examined.</b></p><p><b>Jordan.</b> • <b>Azrak</b>: Sandy desert, 40 km west of Azrak, 17 April 1936, <i>J. E. Dinsmore 11804</i> (E [E 00327349]).</p><p><b>Saudi Arabia.</b> • <b>Turaif</b>: 5 km NNW of Turaif Camp, 22 April 1994, <i>I. S. Collenette 9092</i> (E [E 00092227] & [E 00092228]); near Harrat Al-Harrat Reserve, 04 March 2021, <i>A. Alzahrani 148</i> (MUZ).</p><p><b>Notes.</b></p><p><i>Verbascum transjordanicum</i> is distinguished by its solitary flowers, subsessile glandular stems, simple and sparsely forked hairs on the exterior of the corolla, and five stamens, with the two anterior filaments glabrous near the apex and the three posterior filaments bearing creamy hairs up to the anthers. Phylogenetic analyses (Alzahrani et al. 2024) further support its status as a distinct species.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_18089317 |
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| publishDate | 2025 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Verbascum transjordanicum Murb. Alzahrani, Ali Mohammed Magos Brehm, Joana Ghazanfar, Shahina A. Maxted, Nigel Biodiversity Taxonomy Plantae Tracheophyta Magnoliopsida Lamiales Scrophulariaceae Verbascum Verbascum transjordanicum <p><b>15.</b></p><i>Verbascum transjordanicum</i>Murb. in Lunds Univ. Arsskrift, n. f. xxxv. No. 1 54 (1939)<p>Fig. 28</p><p><b>Type.</b></p><p>Jordan, Sandy desert, 40 km west of Azrak, 17 April 1936, <i>J. E. Dinsmore 11804</i> (isotype E [E 00327349]).</p><p><b>Description.</b></p><p>Annual or biennial herb, pale green, usually branched, many-stemmed from the base, up to 70 cm tall. <i>Indumentum</i> dense glandular hairs with sparse simple and forked hairs above, and dense tomentose with stellate hairs below. <i>Stems</i> erect, robust, terete to angular. <i>Basal leaves</i> rosette, oblong-lanceolate, 2.5–12 × 1–3.5 cm, apex acute, base obtuse, margins crenate to pinnatifid-lobed, lamina darkish green with sparse stellate hairs above and dense white-grey tomentose with stellate hairs below; petiole 1–4 cm. <i>Cauline leaves</i> oblong-lanceolate, 1.6–5 × 1–2 cm, apex acute, base obtuse; sessile. <i>Inflorescence</i> racemose; flowers single in the axil of bracts. <i>Upper bracts</i> ovate-triangular, 1–4 mm, acute. <i>Lower bracts</i> oblong-lanceolate, 4–8 mm, acute. <i>Pedicel</i> covered with dense glandular hairs up to 20 mm long. <i>Bracteoles</i> absent. <i>Calyx</i> 3.4–4.4 mm, lobes oblong-lanceolate, acute, dense glandular. <i>Corolla</i> 20–30 mm across, yellow with dark purple-red hairy blotches around the throat, without pellucid glands, tube up to 1 mm, dense papillose hairs inside, spare glandular, simple, and forked hairs outside. <i>Stamens</i> five, 3–5 mm long. <i>Filaments</i> yellow with creamy hairs, two anterior glabrous near the apex, three posteriors with hairs up to anthers. <i>Anthers</i> all reniform. <i>Ovary</i> globose-ovoid, sparse glandular hairs. <i>Style</i> up to 7 mm long, filiform, green. <i>Stigma</i> capitate. <i>Capsule</i> 4–6 × 3–4 mm, globose-ovoid, sparse glandular hairs. <i>Seeds</i> bothrospermous.</p><p><b>Distribution.</b></p><p>Jordan and Saudi Arabia.</p><p><b>Distribution in the Arabian Peninsula.</b></p><p>It is a native species to Saudi Arabia, which is known from Turaif province (near Harrat Al-Harrat Reserve), northern Saudi Arabia (Fig. 29).</p><p><b>Ecology.</b></p><p><i>Verbascum transjordanicum</i> grows on a limestone plateau with basalt rock at altitudes ranging from 600 to 832 m. Associated plants include <i>Achillea fragrantissima</i> (Forssk.) Sch. Bip., <i>Centaurea sinaica</i> DC., <i>Cornulaca setifera</i> (DC.) Moq., <i>Helianthemum lippii</i> (L.) Dum. Cours., and <i>Hyoscyamus muticus subsp. muticus</i>.</p><p><b>Vernacular name.</b></p><p><i>Transjordan Mullein</i> (English), <i>Abu Ain</i> (Arabic).</p><p><b>Phenology.</b></p><p>Flowering and fruiting from March to June.</p><p><b>Etymology.</b></p><p>The name refers to its occurrence in the Transjordan region, meaning “ across ” or “ beyond ” Jordan.</p><p><b>Specimens examined.</b></p><p><b>Jordan.</b> • <b>Azrak</b>: Sandy desert, 40 km west of Azrak, 17 April 1936, <i>J. E. Dinsmore 11804</i> (E [E 00327349]).</p><p><b>Saudi Arabia.</b> • <b>Turaif</b>: 5 km NNW of Turaif Camp, 22 April 1994, <i>I. S. Collenette 9092</i> (E [E 00092227] & [E 00092228]); near Harrat Al-Harrat Reserve, 04 March 2021, <i>A. Alzahrani 148</i> (MUZ).</p><p><b>Notes.</b></p><p><i>Verbascum transjordanicum</i> is distinguished by its solitary flowers, subsessile glandular stems, simple and sparsely forked hairs on the exterior of the corolla, and five stamens, with the two anterior filaments glabrous near the apex and the three posterior filaments bearing creamy hairs up to the anthers. Phylogenetic analyses (Alzahrani et al. 2024) further support its status as a distinct species.</p> |
| title | Verbascum transjordanicum Murb. |
| topic | Biodiversity Taxonomy Plantae Tracheophyta Magnoliopsida Lamiales Scrophulariaceae Verbascum Verbascum transjordanicum |
| url | https://doi.org/10.5281/zenodo.18089317 |