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Bibliographic Details
Main Authors:	Back, Jinwook, Bang, Hyun Woo
Format:	Recurso digital
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Published:	Zenodo 2026
Subjects:	Biodiversity Taxonomy Animalia Arthropoda Copepoda Harpacticoida Orthopsyllidae Intercristacoxa Intercristacoxa trisetosa
Online Access:	https://doi.org/10.5281/zenodo.18199782
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Intercristacoxa trisetosa sp. nov.Figs 18, 19, 20, 21, 22, 23, 24, 25, 26Type locality.Subtidal, site C (35°55'30.6"N, 129°32'52.3"E, Fig. 1), depth 10 m, sand between cracks in stone, on April 11.Type material.Holotype: • 1 &female; (MABIK CR 00258665), dissected on thirteen slides. Paratypes: • 1 &female; (MABIK CR 00258666), dissected on ten slides; • 1 &male; (MABIK CR 00258667), dissected on nine slides.GenBank accession number.Mitochondrial cytochrome c oxidase subunit I gene (PX 210753); 18 S ribonucleic acid genes (PX 218730 and PX 218731).Description.Female. Total body (Fig. 18 A, B), nearly cylindrical, with minute sensilla dorsally, length from anterior margin of rostrum to posterior margin of caudal rami 1,260 μm, maximum width 240 μm measured at end of cephalothorax; rostrum (Fig. 18 C) well developed, defined at base, tri-angular, with two sensilla; cephalothorax wider than free somites, pleural areas of cephalic shield narrow and posterolateral angles rounded; second and third urosomites (genital double somite) separated dorsally and laterally, fused ventrally but discontinuous internal chitinous rib indicating original segmentation, genital apparatus located at anterior of genital double somite (Fig. 22 A); anal operculum deeply curved (Fig. 18 D).Caudal rami (Figs 22 A, 23 A – C): parallel; each ramus with seven setae; seta V bi-articulated, consists of spine-like seta at top and small pinnate seta attached at end (Fig. 23 B); seta VI bare at inner distal corner; seta VII tri-articulated at base.A 1 (Fig. 19 A): four-segmented; seg – 1 with one thorn-like process at distal outer edge; seg – 3 longest, with one ae; armature formula: 1 – [1], 2 – [7], 3 – [10 + ae], 4 – [12]; apical ae not observed, but three apical setae fused at base.A 2 (Fig. 19 B): three-segmented; coxa bare; allobasis without abexopodal seta; exp one-segmented with two setae; free enp with four spines, three geniculate setae, and one bare seta.Md (Fig. 20 A): gnathobase with one uni-pinnate seta laterally and several teeth; basis of palp with two setae; enp one-segmented with three bare setae; exp incorporated into basis, represented by small protrusion with one bare seta.Mxl (Fig. 20 B): praecoxa trapezoidal; arthrite well developed with two juxtaposed bare setae on anterior surface, two pinnate setae laterally, and six elements around distal margin; coxa with one spine and one seta; basis with one spine and three setae; enp incorporated in basis, represented by small protrusion with three bare setae; exp one-segmented, with two setae.Mxa (Fig. 20 C): syncoxa with three endites; proximal endite vestigial, represented by one seta; second endite with one strong spine and two bare setae; distal endite with one geniculate and two bare setae; allobasis produced into strong curved claw, accessory armature consisting of two setae; and enp represented by three bare setae fused at base.Mxp (Fig. 20 D): three-segmented; syncoxa with one pinnate seta; basis slender, elongate, and bare; enp represented by an acutely recurved claw with two accessory setae.P 1 (Fig. 21 A): coxa with two outer serrate crests; basis with one outer and one inner pinnate setae, ornamented with small frills on base of enp; exp one-segmented, with three outer and two distal setae, ornamented with two small protuberances; enp two-segmented, prehensile; enp – 1 elongate, unarmed; enp – 2 rectangular, with one curved claw and one seta.P 2 (Fig. 21 B): coxa bare; basis with one outer pinnate seta; exp one-segmented, with three spines and two pinnate setae; enp absent.P 3 (Fig. 21 C): coxa bare, basis with one outer bare seta; exp one-segmented with two outer spines and three pinnate setae; enp one-segmented with three setae.P 4 (Fig. 22 C): coxa bare; basis with one outer bare seta; exp one-segmented, with two spines and three pinnate setae, ornamented with a row of setules along inner and outer margins; enp one-segmented, with three setae.Armature formulae as follows:P 5 (Fig. 23 D): basal part with one outer bare seta; endopodal lobe with four pinnate setae, an abnormality observed in innermost seta of paratype (Fig. 23 E, black arrow); exp defined at base with six pinnate setae.P 6 (Fig. 22 B): P 6 vestigial, represented by a small bump with two bare setae.Male. Total body (Fig. 24 A): length from anterior margin of rostrum to posterior margin of caudal rami 945 μm; maximum width 150 μm measured at midway of cephalothorax.General body shape, ornamentation, and sensilla pattern same as in females, except for genital double somites and sexual dimorphism in A 1, P 3, P 4, P 5, and P 6.A 1 (Fig. 25 A – C): six-segmented, robust, chirocer; seg – 1 with one thorn-like process at distal outer edge; seg – 5 swollen, with pad-like process near anterior margin, three processes in middle, and ae fused at base to one bare seta, armature formula: 1 – [1], 2 – [10], 3 – [6], 4 – [2], 5 – [7 + (1 + ae)], 6 – [12], apical ae not observed.P 3 (Fig. 26 A): coxa bare; basis with one outer bare seta; exp one-segmented, with two spines and three setae; enp two-segmented; enp – 1 small, bare; enp – 2 elongated, armed with a spinous apophysis, with two lateral pinnate and one apical minute setae.P 4 (Fig. 26 B): coxa bare, basis with one outer bare seta; exp one-segmented, with two spines and three setae; enp one-segmented, with three setae (middle setae shorter than in females).P 5 (Fig. 26 C): benp represented by small protrusion with two pinnate setae; exp defined at base, with four pinnate setae.P 6 (Fig. 26 D): asymmetrical, with functional right member articulating at base and closing off genital aperture, and left member fused at base to genital somite; each P 6 with two pinnate setae.Etymology.The species name refers to the character of the P 3 and P 4 enp armed with three setae.

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