Epicypta ganengsengi Amorim & Oliveira & Balbi & Ang & Torres & Yeo & Srivathsan & Meier 2025, sp. n.
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| author | Amorim, Dalton De Souza Oliveira, Sarah Siqueira Balbi, Maria Isabel P. A. Ang, Yuchen Torres, Ambrosio Yeo, Darren Srivathsan, Amrita Meier, Rudolf |
| author_facet | Amorim, Dalton De Souza Oliveira, Sarah Siqueira Balbi, Maria Isabel P. A. Ang, Yuchen Torres, Ambrosio Yeo, Darren Srivathsan, Amrita Meier, Rudolf |
| contents | <p><b><i>Epicypta ganengsengi</i> Amorim & Oliveira sp. n. (Figs. 132A–C, 133A, B)</b></p><p>https://singapore.biodiversity.online/species/A-Arth-Hexa-Diptera-000795</p><p>urn:lsid:zoobank.org:act: B9A73545-2327-495E-8937- 10C764228B43</p><p>Diagnosis</p><p>Head brown, antennal scape and pedicel ochre-yellowish, flagellum greyish-brown; scutum dark brown, a diffuse dark ochre-yellowish area along anterior margin, scutellum blackish-brown; pleural sclerites dark brown, antepronotum and proepisternum dark ochre-yellowish. Coxae and femora whitish, on fore leg with brownish light tinge, mid and hind coxae with a brown band on proximal end, tip brownish. Wing membrane light brownish, cells c and br darker. C barely produced beyond tip of R 5; M 1+2 0.58× r-m length. Dorsal macrotrichia on posterior veins M 1 and M 2, M 4, and on CuA distal fourth; macrotrichia on anal lobe membrane. Abdominal tergites 1–6 dark brown, tergite 7 brownish-yellow. Gonocoxites fused medially, with a short and wide medioventral projection, a pair of laterodistal projections over five times longer than length of syngonocoxite medially, with scattered fine setae and a couple of subdistal strong setae and some longer setae at tip. Gonostylus simple, digitiform. Aedeagus rhomboid, bare, well sclerotized, parameres weakly sclerotized, with a pair of strong, straight spines close together medially. Tergite 9 present as a pair of wide posterior lobes covered with microtrichia and setae, straight on outer margins, wider at base. Female terminalia with short incision on posterior margin of sternite 8.</p><p><i>Epicypta_ganengsengi _ ZRCBDP 0133402_hapZRCBDP0132824_SMH_holotype</i> [28: T, 85: G, 133: T, 146: A, 172: C, 193: T, 277: G] tctttcatctactattgctcacgcaggTtcttctgt agatttagctattttttctcttcatttagcaggaatttcttcaattttGggagctattaatt ttattactactattatcaatatacgagcccctggTatttcttttgatAaaatacctttat ttgtttgatcagtCttaattacagctattttactTcttttatctttacctgttttagctgga gctattactatattattaacagatcgaaatttaaatacttctttttttgaccctgcGgga gggggagaccctattttatatcaacatttattt</p><p><b>Description</b></p><p>Male. Wing length, 2.38; width, 0.83. Head. Vertex dark caramel-brown, occiput dark caramel-brown. Scape and pedicel ochre-yellowish, first two flagellomeres light brown, remaining yellowish-brown at basal third, light brown at distal two-thirds. Face and clypeus light caramel-brown. Maxillary palpomeres light brownish. Scape 1.6× pedicel length; flagellomere 4 1.5× longer than wide, covered with scattered whitish setae. Face and clypeus light caramel-brown; face covered with short darker setae, clypeus short, densely covered with setulae. Palpomeres light brownish; labella whitish-yellow. Occiput with three longer setae dorsally to eye anteriorly to lateral ocellus, one dorsally to ocellus, seven posteriorly to ocellus. Scape 1.6× pedicel length, flagellomere 4 1.5× width. Palpomere 4 1.1× palpomere 3 length, palpomere 5 2.3× palpomere 4 length. Thorax. Scutum dark caramel-brown, scutellum blackish-brown. Antepronotum and proepisternum light caramel-brown, anepisternum dark caramel-brown, light along anterior margin, other sclerites greyish-brown, mediotergite dark brown. Pleural membrane ochre-yellow. Haltere ochre-yellowish. Scutum with 5+1 supra-alars (first small) and three pairs of prescutellar bristles, scutellum with two pairs of bristles along posterior margin. Proepisternum with three bristles, anepisternum with five bristles along posterior margin. Mesepimeron with two bristles and 16 small setae, laterotergite with two long setae and seven small setae. Metepisternum with three fine setae. Legs. Coxae light ochre-yellowish, mid and hind coxae with a brown band basally, hind coxa with a brown posterior mark distally; femora light ochre-yellowish with a brown tinge along dorsal and ventral edges; tibiae ochre-yellowish, mid and hind tibiae greyish-brown at basal end, yellowish-brown at tip, tarsi light greyish-brown. Fore tibia with one strong seta on distal third of inner face; mid tibia with three irregular dorsolateral rows of 3–5 small bristles, one strong seta on distal third of inner face; hind tibia with three irregular rows of 3–5 dorsolateral small bristles and five larger setae on distal third of inner face. Fore leg tarsomere 1 1.2× tibia length [tarsomeres 2–5 broken]. Hind tibial inner spur over 4.6× tibia width at apex. Wing (Fig. 132B). Membrane light brown fumose. C not extending beyond apex of R 5. Sc barely produced. R 1 reaching C on distal fifth of wing; R 5 reaching C slightly before level of tip of M 1. First sector of Rs nearly transverse, 0.52× r-m length; r-m almost longitudinal. M 1+2 0.58× r-m length; bM about 4.7× r-m length; first sector of CuA about 0.18× length of second sector of CuA. Cubital pseudovein absent, CuP extending slightly beyond level of origin of M 4. Anal fold gradually curved, nearly reaching wing margin. Posterior veins M 1 and M 2 with dorsal macrotrichia on most of their length, M 4 with macrotrichia on distal 3/4, CuA with macrotrichia on distal fourth, macrotrichia on anal lobe membrane. Abdomen. Abdominal tergites 1–6 brown, posterior margin of tergite 6 yellowish, tergite 7 yellowish; sternites 1–7 whitish. Sternite 2 with a pair of strong brown bristles. Terminalia (Fig. 133A, B). Gonocoxites fused medially, no suture present, bare ventrally, a large and short medioventral projection, a laterodistal projection dorsally to insertion of gonostylus at each side that is over five times longer than length of syngonocoxite medially, with scattered fine setae along entire length, a couple of subdistal strong setae and some longer setae at tip. Gonostylus simple, digitiform, with fine setae, slightly stronger distally. Aedeagus rhomboid, bare, well sclerotized, parameres weakly sclerotized, projecting posteriorly beyond aedeagus, with a pair of strong, straight spines close together medially. Gonocoxal bridge wide, apodemes as a pair of long slender sclerotized bands. Tergite 9 present as a pair of wide posterior lobes covered with microtrichia and setae, straight on outer margins, wider at base. Cerci not recognizable.</p><p>Female (Fig. 132A). As male, except for the following. Terminalia (Fig. 132C). Light brownish-yellow, cerci lighter. Sternite 8 trapezoid, elongate, posterior margin straight, with a rather deep medial incision, widespread microtrichia and small setae, posterior margin with longer setae. Sternite 9 with anterior apodeme extending beyond anterior end of terminalia, widening close to tip, genital chamber bell-shaped, lateral arms wide, distal end triangular, acute, bare. Tergite 8 wide, lateral lobes projected posteriorly with setae, with a slender medial connection. Tergite 9+10 present as a pair of slender sclerotized inclined band, connected to sternite 9 lateroventrally. Sternite 10 triangular, with some setae close to tip directed dorsally. Cercus large, cercomeres 1 and 2 fused, no sign of suture, basal 3/4 wider, posterior end digitiform, distal setae longer.</p><p>Material examined</p><p>Holotype: male, ZRCBDP 0133402, National University of Singapore (Uhall), 17.may.17, MIP leg. (slide-mounted).</p><p>Paratypes (5 females): ZRCBDP 0049044, Nee Soon (NS2), 07.jan.15, MIP leg.; ZRCBDP 0049300, National University of Singapore (Icube), 01.apr.15, MIP leg. (website photo specimen, slide-mounted); ZRCBDP 0072662, Bukit Timah, primary forest (BT05), 28.dec.16, MIP leg.; ZRCBDP 0072722, Bukit Timah, old secondary forest (BT01), 14.dec.16, MIP leg. (slide-mounted); ZRCBDP 0279158, Singapore, 7.jun.18, MIP leg. (slide-mounted).</p><p>Additional sequenced specimens: ZRCBDP 0132824, National University of Singapore (PGP), urban forest, 24.may.17; ZRCBDP 0133115, National University of Singapore (PGP), urban forest, 17.may.17; ZRCBDP 0133129, National University of Singapore (PGP), urban forest, 17.may.17; ZRCBDP 0133169, National University of Singapore (Uhall), urban forest, 26.apr.17, MIP leg.; ZRCBDP 0133378, National University of Singapore (PGP), urban forest, 14.may.17; ZRCBDP 0133417, National University of Singapore (PGP), urban forest, 21.may.17; ZRCBDP 0133490, National University of Singapore (PGP), urban forest, 31.may.17; ZRCBDP 0133504, Singapore, date range 2012–2018, MIP leg.; ZRCBDP 0133552; ZRCBDP 0279142, Singapore, date range 2012–2018, MIP leg.; ZRCBDP 0279159, Singapore, date range 2012–2018, MIP leg.; ZRCBDP 0284194, Kranji Marshes (KM03), freshwater swamp, date range 2018-2019, MIP leg; ZRCBDP 0133158, National University of Singapore (RVR), 03.may.17; ZRCBDP 0133945, Singapore.</p><p>Etymology</p><p>The species epithet honors GAN ENG SENG (1844–1899), a businessman, philanthropist, and early pioneer in Singapore. He is known for his generosity to many charitable causes in Malaya and Singapore during the British colonial era. In 1885, he founded the first school established by overseas Chinese in Singapore, one of the oldest educational institutions in the nation-state.</p><p><b>Remarks</b></p><p>There are five haplotypes for <i>Epicypta ganengsengi</i> <b>sp. n.</b>, clustering together into one mOTU using all species delimitation methods. Specimens were collected in both the urban and swamp forests.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_18313195 |
| institution | Zenodo |
| language | |
| publishDate | 2025 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Epicypta ganengsengi Amorim & Oliveira & Balbi & Ang & Torres & Yeo & Srivathsan & Meier 2025, sp. n. Amorim, Dalton De Souza Oliveira, Sarah Siqueira Balbi, Maria Isabel P. A. Ang, Yuchen Torres, Ambrosio Yeo, Darren Srivathsan, Amrita Meier, Rudolf Biodiversity Taxonomy Animalia Arthropoda Insecta Diptera Mycetophilidae Epicypta Epicypta ganengsengi <p><b><i>Epicypta ganengsengi</i> Amorim & Oliveira sp. n. (Figs. 132A–C, 133A, B)</b></p><p>https://singapore.biodiversity.online/species/A-Arth-Hexa-Diptera-000795</p><p>urn:lsid:zoobank.org:act: B9A73545-2327-495E-8937- 10C764228B43</p><p>Diagnosis</p><p>Head brown, antennal scape and pedicel ochre-yellowish, flagellum greyish-brown; scutum dark brown, a diffuse dark ochre-yellowish area along anterior margin, scutellum blackish-brown; pleural sclerites dark brown, antepronotum and proepisternum dark ochre-yellowish. Coxae and femora whitish, on fore leg with brownish light tinge, mid and hind coxae with a brown band on proximal end, tip brownish. Wing membrane light brownish, cells c and br darker. C barely produced beyond tip of R 5; M 1+2 0.58× r-m length. Dorsal macrotrichia on posterior veins M 1 and M 2, M 4, and on CuA distal fourth; macrotrichia on anal lobe membrane. Abdominal tergites 1–6 dark brown, tergite 7 brownish-yellow. Gonocoxites fused medially, with a short and wide medioventral projection, a pair of laterodistal projections over five times longer than length of syngonocoxite medially, with scattered fine setae and a couple of subdistal strong setae and some longer setae at tip. Gonostylus simple, digitiform. Aedeagus rhomboid, bare, well sclerotized, parameres weakly sclerotized, with a pair of strong, straight spines close together medially. Tergite 9 present as a pair of wide posterior lobes covered with microtrichia and setae, straight on outer margins, wider at base. Female terminalia with short incision on posterior margin of sternite 8.</p><p><i>Epicypta_ganengsengi _ ZRCBDP 0133402_hapZRCBDP0132824_SMH_holotype</i> [28: T, 85: G, 133: T, 146: A, 172: C, 193: T, 277: G] tctttcatctactattgctcacgcaggTtcttctgt agatttagctattttttctcttcatttagcaggaatttcttcaattttGggagctattaatt ttattactactattatcaatatacgagcccctggTatttcttttgatAaaatacctttat ttgtttgatcagtCttaattacagctattttactTcttttatctttacctgttttagctgga gctattactatattattaacagatcgaaatttaaatacttctttttttgaccctgcGgga gggggagaccctattttatatcaacatttattt</p><p><b>Description</b></p><p>Male. Wing length, 2.38; width, 0.83. Head. Vertex dark caramel-brown, occiput dark caramel-brown. Scape and pedicel ochre-yellowish, first two flagellomeres light brown, remaining yellowish-brown at basal third, light brown at distal two-thirds. Face and clypeus light caramel-brown. Maxillary palpomeres light brownish. Scape 1.6× pedicel length; flagellomere 4 1.5× longer than wide, covered with scattered whitish setae. Face and clypeus light caramel-brown; face covered with short darker setae, clypeus short, densely covered with setulae. Palpomeres light brownish; labella whitish-yellow. Occiput with three longer setae dorsally to eye anteriorly to lateral ocellus, one dorsally to ocellus, seven posteriorly to ocellus. Scape 1.6× pedicel length, flagellomere 4 1.5× width. Palpomere 4 1.1× palpomere 3 length, palpomere 5 2.3× palpomere 4 length. Thorax. Scutum dark caramel-brown, scutellum blackish-brown. Antepronotum and proepisternum light caramel-brown, anepisternum dark caramel-brown, light along anterior margin, other sclerites greyish-brown, mediotergite dark brown. Pleural membrane ochre-yellow. Haltere ochre-yellowish. Scutum with 5+1 supra-alars (first small) and three pairs of prescutellar bristles, scutellum with two pairs of bristles along posterior margin. Proepisternum with three bristles, anepisternum with five bristles along posterior margin. Mesepimeron with two bristles and 16 small setae, laterotergite with two long setae and seven small setae. Metepisternum with three fine setae. Legs. Coxae light ochre-yellowish, mid and hind coxae with a brown band basally, hind coxa with a brown posterior mark distally; femora light ochre-yellowish with a brown tinge along dorsal and ventral edges; tibiae ochre-yellowish, mid and hind tibiae greyish-brown at basal end, yellowish-brown at tip, tarsi light greyish-brown. Fore tibia with one strong seta on distal third of inner face; mid tibia with three irregular dorsolateral rows of 3–5 small bristles, one strong seta on distal third of inner face; hind tibia with three irregular rows of 3–5 dorsolateral small bristles and five larger setae on distal third of inner face. Fore leg tarsomere 1 1.2× tibia length [tarsomeres 2–5 broken]. Hind tibial inner spur over 4.6× tibia width at apex. Wing (Fig. 132B). Membrane light brown fumose. C not extending beyond apex of R 5. Sc barely produced. R 1 reaching C on distal fifth of wing; R 5 reaching C slightly before level of tip of M 1. First sector of Rs nearly transverse, 0.52× r-m length; r-m almost longitudinal. M 1+2 0.58× r-m length; bM about 4.7× r-m length; first sector of CuA about 0.18× length of second sector of CuA. Cubital pseudovein absent, CuP extending slightly beyond level of origin of M 4. Anal fold gradually curved, nearly reaching wing margin. Posterior veins M 1 and M 2 with dorsal macrotrichia on most of their length, M 4 with macrotrichia on distal 3/4, CuA with macrotrichia on distal fourth, macrotrichia on anal lobe membrane. Abdomen. Abdominal tergites 1–6 brown, posterior margin of tergite 6 yellowish, tergite 7 yellowish; sternites 1–7 whitish. Sternite 2 with a pair of strong brown bristles. Terminalia (Fig. 133A, B). Gonocoxites fused medially, no suture present, bare ventrally, a large and short medioventral projection, a laterodistal projection dorsally to insertion of gonostylus at each side that is over five times longer than length of syngonocoxite medially, with scattered fine setae along entire length, a couple of subdistal strong setae and some longer setae at tip. Gonostylus simple, digitiform, with fine setae, slightly stronger distally. Aedeagus rhomboid, bare, well sclerotized, parameres weakly sclerotized, projecting posteriorly beyond aedeagus, with a pair of strong, straight spines close together medially. Gonocoxal bridge wide, apodemes as a pair of long slender sclerotized bands. Tergite 9 present as a pair of wide posterior lobes covered with microtrichia and setae, straight on outer margins, wider at base. Cerci not recognizable.</p><p>Female (Fig. 132A). As male, except for the following. Terminalia (Fig. 132C). Light brownish-yellow, cerci lighter. Sternite 8 trapezoid, elongate, posterior margin straight, with a rather deep medial incision, widespread microtrichia and small setae, posterior margin with longer setae. Sternite 9 with anterior apodeme extending beyond anterior end of terminalia, widening close to tip, genital chamber bell-shaped, lateral arms wide, distal end triangular, acute, bare. Tergite 8 wide, lateral lobes projected posteriorly with setae, with a slender medial connection. Tergite 9+10 present as a pair of slender sclerotized inclined band, connected to sternite 9 lateroventrally. Sternite 10 triangular, with some setae close to tip directed dorsally. Cercus large, cercomeres 1 and 2 fused, no sign of suture, basal 3/4 wider, posterior end digitiform, distal setae longer.</p><p>Material examined</p><p>Holotype: male, ZRCBDP 0133402, National University of Singapore (Uhall), 17.may.17, MIP leg. (slide-mounted).</p><p>Paratypes (5 females): ZRCBDP 0049044, Nee Soon (NS2), 07.jan.15, MIP leg.; ZRCBDP 0049300, National University of Singapore (Icube), 01.apr.15, MIP leg. (website photo specimen, slide-mounted); ZRCBDP 0072662, Bukit Timah, primary forest (BT05), 28.dec.16, MIP leg.; ZRCBDP 0072722, Bukit Timah, old secondary forest (BT01), 14.dec.16, MIP leg. (slide-mounted); ZRCBDP 0279158, Singapore, 7.jun.18, MIP leg. (slide-mounted).</p><p>Additional sequenced specimens: ZRCBDP 0132824, National University of Singapore (PGP), urban forest, 24.may.17; ZRCBDP 0133115, National University of Singapore (PGP), urban forest, 17.may.17; ZRCBDP 0133129, National University of Singapore (PGP), urban forest, 17.may.17; ZRCBDP 0133169, National University of Singapore (Uhall), urban forest, 26.apr.17, MIP leg.; ZRCBDP 0133378, National University of Singapore (PGP), urban forest, 14.may.17; ZRCBDP 0133417, National University of Singapore (PGP), urban forest, 21.may.17; ZRCBDP 0133490, National University of Singapore (PGP), urban forest, 31.may.17; ZRCBDP 0133504, Singapore, date range 2012–2018, MIP leg.; ZRCBDP 0133552; ZRCBDP 0279142, Singapore, date range 2012–2018, MIP leg.; ZRCBDP 0279159, Singapore, date range 2012–2018, MIP leg.; ZRCBDP 0284194, Kranji Marshes (KM03), freshwater swamp, date range 2018-2019, MIP leg; ZRCBDP 0133158, National University of Singapore (RVR), 03.may.17; ZRCBDP 0133945, Singapore.</p><p>Etymology</p><p>The species epithet honors GAN ENG SENG (1844–1899), a businessman, philanthropist, and early pioneer in Singapore. He is known for his generosity to many charitable causes in Malaya and Singapore during the British colonial era. In 1885, he founded the first school established by overseas Chinese in Singapore, one of the oldest educational institutions in the nation-state.</p><p><b>Remarks</b></p><p>There are five haplotypes for <i>Epicypta ganengsengi</i> <b>sp. n.</b>, clustering together into one mOTU using all species delimitation methods. Specimens were collected in both the urban and swamp forests.</p> |
| title | Epicypta ganengsengi Amorim & Oliveira & Balbi & Ang & Torres & Yeo & Srivathsan & Meier 2025, sp. n. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Diptera Mycetophilidae Epicypta Epicypta ganengsengi |
| url | https://doi.org/10.5281/zenodo.18313195 |