Epicypta wallacei Amorim & Oliveira & Balbi & Ang & Torres & Yeo & Srivathsan & Meier 2025, sp. n.
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| author | Amorim, Dalton De Souza Oliveira, Sarah Siqueira Balbi, Maria Isabel P. A. Ang, Yuchen Torres, Ambrosio Yeo, Darren Srivathsan, Amrita Meier, Rudolf |
| author_facet | Amorim, Dalton De Souza Oliveira, Sarah Siqueira Balbi, Maria Isabel P. A. Ang, Yuchen Torres, Ambrosio Yeo, Darren Srivathsan, Amrita Meier, Rudolf |
| contents | <p><i>Epicypta wallacei</i> Amorim & Oliveira sp. n.</p><p>(Figs. 143A–D, 144A, B)</p><p>https://singapore.biodiversity.online/species/A-Arth-Hexa-Diptera-000738</p><p>urn:lsid:zoobank.org:act: 724B5B7C-E6EC-4285-9027- 260BCDA8A75A</p><p><b>Diagnosis</b></p><p>Head and scutum ochre-yellowish, antenna brownish; scutellum yellowish-brown; pleural sclerites dark ochre-yellowish, antepronotum and proepisternum lighter, laterotergite, metepisternum, and mediotergite brown. Coxae and femora whitish, a small brown mark at basal tip of hind coxa and at tip of hind femur. Wing membrane light brownish, cell c and cell br darker. C not produced beyond tip of R 5; Sc long; M 1+2 about half of r-m length. Dorsal macrotrichia on posterior veins M 1 and M 2, M 4, and on CuA close to tip, macrotrichia on anal lobe membrane. Abdominal tergites 1–6 greyish-brown with light ochre-yellow anterolateral corners, wider on tergites 4–5, tergite 7 ochre-yellow, terminalia yellowish. Male terminalia gonocoxite with a medioventral digitiform projection and a digitiform laterodistal extension reaching way beyond tip of gonostylus; gonostylus simple, elongate, dorsoventrally compressed; aedeagus with a pair of separate tubular extensions each with a gonopore; parameres triangular with a pair of strong, curved spines close to tip; tergite 9 present as a pair of long laterodistal separate extensions, much longer than gonostylus or gonocoxite lobes.</p><p><i>Epicypta_wallacei _ ZRCBDP 0048465_hapZRCBDP0048465_SMH_holotype</i> [1: C, 7: C, 19: C, 29: T, 67: C, 106: C, 178: C, 196: T, 199: T] CctatcCtcaactattgcCcatg ctggaTcttctgtagatttagctattttttctttacatttagcCggaatctcctcaatttt aggggcaattaattttattacCacaattattaatatacgatccccaggaattacttttgatcgaatacctctatttgtttgatcagtattaatCacagctattttacttctTctTtcttt acctgttttagcgggagctattacaatattattaacagatcgaaatttaaatacttcttt ttttgatccagcgggaggaggggatcctattttatatcaacatttattt</p><p><b>Description</b></p><p>Male (Fig. 143A). Wing length, 2.21; width, 0.80. Head. Head ochre-yellowish. Scape greyish-yellow, pedicel ochre-yellowish, flagellomeres light brown. Face and clypeus ochre-yellowish, palpomeres 1–3 ochre-yellowish, palpomeres 4–5 whitish-yellow, labella cream-yellow. Occiput with four longer setae dorsally to eye anteriorly to lateral ocellus, two dorsally to ocellus, eight posteriorly to ocellus. Scape about 1.3× pedicel length; flagellomere 4 1.7× as long as wide. Palpomere 4 1.1× palpomere 3 length, palpomere 5 2.0× palpomere 4 length. Thorax. Scutum ochre-yellow, scutellum yellowish-brown with lighter corners. Pleural sclerites light ochre-yellowish, laterotergite and metepisternum greyish-brown, mediotergite yellowish-brown. Haltere pedicel whitish, knob brownish basally, whitish distally. Pleural membrane yellowish. Scutum with seven supra-alar bristles and three pairs of prescutellar bristles, plus a pair of long setae between inner two, three pairs of scutellar bristles and one outer pair of longer setae. Proepisternum with three bristles, anepisternum with five bristles along posterior margin. Mesepimeron with three bristles and 20 small setae, laterotergite with three bristles and 18 smaller setae. Metepisternum with 29 small setae. Legs. Fore coxa whitish-yellow with a light brownish tinge, mid and hind coxae whitish, hind coxa with a greyish-brown mark at basal end; femora, tibiae, and tarsi light whitish-yellow, tarsi darker, hind femur with brownish mark at distal end. Fore tibia with two dorsolateral bristles, mid tibia with two dorsolateral rows of 3–5 bristles and three bristles along ventral edge, hind tibia with a row of 5–6 bristles dorsolaterally, a line of five long setae on outer lateral face. Fore leg tarsomere 1 1.1× tibia length, 1.7× tarsomere 2 length, mid and hind tarsomeres 1–3 with rows of ventral setae besides trichia. Hind tibia inner spur 5.0× tibia width at apex. Wing (Fig. 143B). Membrane fumose light brown, darker on cells c, br and r1. Sc barely produced. C barely extending beyond apex of R 5. R 1 reaching C on distal sixth, R 5 reaching C slightly before level of tip of M 1. First sector of Rs oblique, slightly over half of r-m length; r-m almost longitudinal. M 1+2 0.8× r-m length; bM 5.0× r-m length. First sector of CuA 0.45× length of second sector of CuA. Cubital pseudovein barely reaching level of origin of M 4. Anal fold long, only gently curved, almost reaching wing margin. Posterior veins M 1, M 2, and M 4 with dorsal macrotrichia along more than half their length, CuA with macrotrichia on distal fourth of wing; macrotrichia present on anal lobe membrane. Abdomen. Abdominal tergite 1 greyish-brown with dark brown mark along posterior margin, tergites 2–6 greyish-brown medially, cream-yellow laterally and along anterior margin, cream-yellow marks larger on segments 4–5, tergite 7 cream-yellow; sternites 1–7 whitish-yellow. Sternite 2 with a pair of strong brown bristles. Terminalia (Fig. 143C, D). Whitish-yellow. Gonocoxites fused medially, suture present, bare ventrally, a medioventral digitiform projection with setulae at tip sided by a pair of short pointed lobes, each with a small seta at tip, a pair of long, digitiform laterodistal extensions reaching way beyond tip of gonostylus. Gonostylus simple, elongate, dorsoventrally compressed, with setae along inner margin and at distal half on both faces. Aedeagus with an anterior medial ejaculatory apodeme, widening midway to apex, then divided distally into a pair of separate tubular extensions with independent gonopores, laterally with a pair of long, slender distal projections, slightly capitate at tip. Parameres as a triangular sclerite dorsal to aedeagus extending distally, with a medial short beak at tip, a pair of small setae close to each other subapically, a pair of strong, curved spines close to tip. Gonocoxal bridge present, slender medially, no apodemes visible. Tergite 9 present as a pair of long laterodistal extensions, much longer than gonostylus or gonocoxite extensions, setose at tip. Cerci not visible.</p><p>Female. As male, except for the following. Wing length, 2.27; width, 0.86. Terminalia (Fig. 144A, B). Brownish-yellow. Sternite 8 wide anteriorly, subquadrate, posterior margin slightly more slender than base, a very short medial incision, covered with microtrichia and fine setae, longer setae medially. Sternite 9 wide medially, anterior end slender, extending beyond anterior end of sternite 8, gonopore at center of a well-sclerotized plate. Tergite 8 wide, short medially, connecting a pair of lateral short lobes with fine setae, partially overlapping to sternite 8. Tergite 9+10 short, slender, bare. Cercomeres 1 and 2 apparently fused, no sign of suture, elongate, distal end curved ventrally, covered with microtrichia and setae, setae at tip longer.</p><p>Material examined</p><p>Holotype: male, ZRCBDP 0048465, Nee Soon (NS1), swamp forest, 04.apr.12, MIP leg. (website photo specimen, slide-mounted).</p><p>Paratypes (4 females): ZRCBDP 0048832, Nee Soon (NS1), 14.jan.15, MIP leg. (slide-mounted); ZRCBDP 0048868, Nee Soon (NS1), 31.dec.14, MIP leg. (slide-mounted); ZRCBDP 0048884, Nee Soon (NS1), 31.dec.14, MIP leg.; ZRCBDP 0133462, Singapore [2012-2018], MIP leg.</p><p>Etymology</p><p>The species epithet honors ALFRED RUSSEL WALLACE (1823– 1913), a British naturalist, biogeographer, anthropologist, evolutionist, and illustrator. He is best known for independently originating the theory of evolution by natural selection and recognizing phylogenetic relationships among species. His 1858 paper on the subject was one of the triggers for CHARLES DARWIN to publish his own writings on evolution. Aside from his scientific work, he was a social activist, critical of the unjust social and economic system in 19th-century Britain. He carried out extensive field work in the Amazon and in the Malay Archipelago, including Singapore.</p><p><b>Remarks</b></p><p>There are three haplotypes of <i>Epicypta wallacei</i> <b>sp. n.</b>, which cluster together based on all species delimitation methods.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_18313242 |
| institution | Zenodo |
| language | |
| publishDate | 2025 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Epicypta wallacei Amorim & Oliveira & Balbi & Ang & Torres & Yeo & Srivathsan & Meier 2025, sp. n. Amorim, Dalton De Souza Oliveira, Sarah Siqueira Balbi, Maria Isabel P. A. Ang, Yuchen Torres, Ambrosio Yeo, Darren Srivathsan, Amrita Meier, Rudolf Biodiversity Taxonomy Animalia Arthropoda Insecta Diptera Mycetophilidae Epicypta Epicypta wallacei <p><i>Epicypta wallacei</i> Amorim & Oliveira sp. n.</p><p>(Figs. 143A–D, 144A, B)</p><p>https://singapore.biodiversity.online/species/A-Arth-Hexa-Diptera-000738</p><p>urn:lsid:zoobank.org:act: 724B5B7C-E6EC-4285-9027- 260BCDA8A75A</p><p><b>Diagnosis</b></p><p>Head and scutum ochre-yellowish, antenna brownish; scutellum yellowish-brown; pleural sclerites dark ochre-yellowish, antepronotum and proepisternum lighter, laterotergite, metepisternum, and mediotergite brown. Coxae and femora whitish, a small brown mark at basal tip of hind coxa and at tip of hind femur. Wing membrane light brownish, cell c and cell br darker. C not produced beyond tip of R 5; Sc long; M 1+2 about half of r-m length. Dorsal macrotrichia on posterior veins M 1 and M 2, M 4, and on CuA close to tip, macrotrichia on anal lobe membrane. Abdominal tergites 1–6 greyish-brown with light ochre-yellow anterolateral corners, wider on tergites 4–5, tergite 7 ochre-yellow, terminalia yellowish. Male terminalia gonocoxite with a medioventral digitiform projection and a digitiform laterodistal extension reaching way beyond tip of gonostylus; gonostylus simple, elongate, dorsoventrally compressed; aedeagus with a pair of separate tubular extensions each with a gonopore; parameres triangular with a pair of strong, curved spines close to tip; tergite 9 present as a pair of long laterodistal separate extensions, much longer than gonostylus or gonocoxite lobes.</p><p><i>Epicypta_wallacei _ ZRCBDP 0048465_hapZRCBDP0048465_SMH_holotype</i> [1: C, 7: C, 19: C, 29: T, 67: C, 106: C, 178: C, 196: T, 199: T] CctatcCtcaactattgcCcatg ctggaTcttctgtagatttagctattttttctttacatttagcCggaatctcctcaatttt aggggcaattaattttattacCacaattattaatatacgatccccaggaattacttttgatcgaatacctctatttgtttgatcagtattaatCacagctattttacttctTctTtcttt acctgttttagcgggagctattacaatattattaacagatcgaaatttaaatacttcttt ttttgatccagcgggaggaggggatcctattttatatcaacatttattt</p><p><b>Description</b></p><p>Male (Fig. 143A). Wing length, 2.21; width, 0.80. Head. Head ochre-yellowish. Scape greyish-yellow, pedicel ochre-yellowish, flagellomeres light brown. Face and clypeus ochre-yellowish, palpomeres 1–3 ochre-yellowish, palpomeres 4–5 whitish-yellow, labella cream-yellow. Occiput with four longer setae dorsally to eye anteriorly to lateral ocellus, two dorsally to ocellus, eight posteriorly to ocellus. Scape about 1.3× pedicel length; flagellomere 4 1.7× as long as wide. Palpomere 4 1.1× palpomere 3 length, palpomere 5 2.0× palpomere 4 length. Thorax. Scutum ochre-yellow, scutellum yellowish-brown with lighter corners. Pleural sclerites light ochre-yellowish, laterotergite and metepisternum greyish-brown, mediotergite yellowish-brown. Haltere pedicel whitish, knob brownish basally, whitish distally. Pleural membrane yellowish. Scutum with seven supra-alar bristles and three pairs of prescutellar bristles, plus a pair of long setae between inner two, three pairs of scutellar bristles and one outer pair of longer setae. Proepisternum with three bristles, anepisternum with five bristles along posterior margin. Mesepimeron with three bristles and 20 small setae, laterotergite with three bristles and 18 smaller setae. Metepisternum with 29 small setae. Legs. Fore coxa whitish-yellow with a light brownish tinge, mid and hind coxae whitish, hind coxa with a greyish-brown mark at basal end; femora, tibiae, and tarsi light whitish-yellow, tarsi darker, hind femur with brownish mark at distal end. Fore tibia with two dorsolateral bristles, mid tibia with two dorsolateral rows of 3–5 bristles and three bristles along ventral edge, hind tibia with a row of 5–6 bristles dorsolaterally, a line of five long setae on outer lateral face. Fore leg tarsomere 1 1.1× tibia length, 1.7× tarsomere 2 length, mid and hind tarsomeres 1–3 with rows of ventral setae besides trichia. Hind tibia inner spur 5.0× tibia width at apex. Wing (Fig. 143B). Membrane fumose light brown, darker on cells c, br and r1. Sc barely produced. C barely extending beyond apex of R 5. R 1 reaching C on distal sixth, R 5 reaching C slightly before level of tip of M 1. First sector of Rs oblique, slightly over half of r-m length; r-m almost longitudinal. M 1+2 0.8× r-m length; bM 5.0× r-m length. First sector of CuA 0.45× length of second sector of CuA. Cubital pseudovein barely reaching level of origin of M 4. Anal fold long, only gently curved, almost reaching wing margin. Posterior veins M 1, M 2, and M 4 with dorsal macrotrichia along more than half their length, CuA with macrotrichia on distal fourth of wing; macrotrichia present on anal lobe membrane. Abdomen. Abdominal tergite 1 greyish-brown with dark brown mark along posterior margin, tergites 2–6 greyish-brown medially, cream-yellow laterally and along anterior margin, cream-yellow marks larger on segments 4–5, tergite 7 cream-yellow; sternites 1–7 whitish-yellow. Sternite 2 with a pair of strong brown bristles. Terminalia (Fig. 143C, D). Whitish-yellow. Gonocoxites fused medially, suture present, bare ventrally, a medioventral digitiform projection with setulae at tip sided by a pair of short pointed lobes, each with a small seta at tip, a pair of long, digitiform laterodistal extensions reaching way beyond tip of gonostylus. Gonostylus simple, elongate, dorsoventrally compressed, with setae along inner margin and at distal half on both faces. Aedeagus with an anterior medial ejaculatory apodeme, widening midway to apex, then divided distally into a pair of separate tubular extensions with independent gonopores, laterally with a pair of long, slender distal projections, slightly capitate at tip. Parameres as a triangular sclerite dorsal to aedeagus extending distally, with a medial short beak at tip, a pair of small setae close to each other subapically, a pair of strong, curved spines close to tip. Gonocoxal bridge present, slender medially, no apodemes visible. Tergite 9 present as a pair of long laterodistal extensions, much longer than gonostylus or gonocoxite extensions, setose at tip. Cerci not visible.</p><p>Female. As male, except for the following. Wing length, 2.27; width, 0.86. Terminalia (Fig. 144A, B). Brownish-yellow. Sternite 8 wide anteriorly, subquadrate, posterior margin slightly more slender than base, a very short medial incision, covered with microtrichia and fine setae, longer setae medially. Sternite 9 wide medially, anterior end slender, extending beyond anterior end of sternite 8, gonopore at center of a well-sclerotized plate. Tergite 8 wide, short medially, connecting a pair of lateral short lobes with fine setae, partially overlapping to sternite 8. Tergite 9+10 short, slender, bare. Cercomeres 1 and 2 apparently fused, no sign of suture, elongate, distal end curved ventrally, covered with microtrichia and setae, setae at tip longer.</p><p>Material examined</p><p>Holotype: male, ZRCBDP 0048465, Nee Soon (NS1), swamp forest, 04.apr.12, MIP leg. (website photo specimen, slide-mounted).</p><p>Paratypes (4 females): ZRCBDP 0048832, Nee Soon (NS1), 14.jan.15, MIP leg. (slide-mounted); ZRCBDP 0048868, Nee Soon (NS1), 31.dec.14, MIP leg. (slide-mounted); ZRCBDP 0048884, Nee Soon (NS1), 31.dec.14, MIP leg.; ZRCBDP 0133462, Singapore [2012-2018], MIP leg.</p><p>Etymology</p><p>The species epithet honors ALFRED RUSSEL WALLACE (1823– 1913), a British naturalist, biogeographer, anthropologist, evolutionist, and illustrator. He is best known for independently originating the theory of evolution by natural selection and recognizing phylogenetic relationships among species. His 1858 paper on the subject was one of the triggers for CHARLES DARWIN to publish his own writings on evolution. Aside from his scientific work, he was a social activist, critical of the unjust social and economic system in 19th-century Britain. He carried out extensive field work in the Amazon and in the Malay Archipelago, including Singapore.</p><p><b>Remarks</b></p><p>There are three haplotypes of <i>Epicypta wallacei</i> <b>sp. n.</b>, which cluster together based on all species delimitation methods.</p> |
| title | Epicypta wallacei Amorim & Oliveira & Balbi & Ang & Torres & Yeo & Srivathsan & Meier 2025, sp. n. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Diptera Mycetophilidae Epicypta Epicypta wallacei |
| url | https://doi.org/10.5281/zenodo.18313242 |