Osmyloberotha dispersa Li, Zhuo, Wang, Nakamine, Yamamoto, Zhang, Ling, Ohl, Aspöck, Aspöck & Liu, 2023, sp. nov.
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2023
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| author | Li, Hong-Yu Zhuo, De Wang, Bo Nakamine, Hiroshi Yamamoto, Shûhei Zhang, Wei-Wei Ling, Jia-Ni Ohl, Michael Aspöck, Ulrike Aspöck, Horst Liu, Xing-Yue |
| author_facet | Li, Hong-Yu Zhuo, De Wang, Bo Nakamine, Hiroshi Yamamoto, Shûhei Zhang, Wei-Wei Ling, Jia-Ni Ohl, Michael Aspöck, Ulrike Aspöck, Horst Liu, Xing-Yue |
| contents | <p><b><i>Osmyloberotha dispersa</i> sp. nov.</b></p><p>(Figs 1, 2)</p><p><b>Material.</b> Holotype, ♂, NIGP171721, amber piece polished in the form of a rectangular, transparent cabochon, with length × width about 13.95 × 7.97 mm, height about 4.86 mm (NIGP). Paratype, 1♂, CAM BA-0016, amber piece also preserving a specimen of <i>Jersiberotha</i> sp., three flies, and a beetle; it is polished in the form of a rectangular, transparent cabochon, with length × width about 34.72 × 10.71 mm, height about 5.52 mm (CAM).</p><p><b>Etymology.</b> The specific name “ <i>dispersa</i> ” (Latin, meaning “dispersed”, “scattered”) refers to the evenly dispersed, irregularly shaped, dark markings on the forewing in the new species.</p><p><b>Diagnosis.</b> Forewing with many evenly dispersed, irregular and dark markings. Hind wing immaculate, two incomplete gradates series. Male tergum 9 narrower than ectoprocts; gonocoxites 9 paired, large, subtriangular in lateral view, distally simple and obtuse.</p><p><b>Locality and horizon.</b> Hukawng Valley in Tanai Township, Myitkyina District of Kachin State, Myanmar, Lowermost Cenomanian.</p><p><b>Description.</b> <i>Measurements and proportions.</i> Male. Body length: 2.50–3.34; LP (0.30): WP (0.25) = 1.20; LP: LPT (0.78) = 0.38; LPf (0.45): LPt (0.48) = 0.94; LFW (4.72): WFW (1.60) = 2.95; LHW (4.18): WHW (1.41) = 2.96.</p><p>Male. Head bearing long and soft setae. Vertex moderately domed, bearing markedly long setae. Postocular region markedly narrow in lateral view. Antenna:scape nearly cylindrical,distally slightly swollen, bearing sparse, long setae, 1.5 times as long as wide, and 1.5 times as long as pedicel; pedicel nearly as long as wide, and slightly longer than flagellomere 1; flagellum filiform, with about 33 segments; each flagellomere cylindrical, nearly as long as wide; terminal flagellomere conical, apically tapering into a point. Terminal maxillary and labial palpomeres each proximally distinctly swollen and tapering to an acute point.</p><p>Thorax bearing markedly long setae. Pronotum short, nearly 1/3 length of pterothorax. Forelegs: profemur nearly as long as protibia; protarsus 5-segmented: tarsomere 1 nearly as long as tarsomeres 2–3 combined, tarsomeres 2–3, 5 subequal, tarsomere 4 slightly shorter than tarsomere 3; tarsomeres 1–4 each distally bearing two pairs of thickened setae (spurs). Mid and hind legs similar to forelegs.</p><p>Wings acute, subfalcate, branches of RP mostly distally bifurcate or simple, some trifurcate. Forewing with many evenly dispersed, irregular, dark markings; costal space narrow, not dilated; costal vein distally separate from ScP; ScA absent; recurrent humeral crossvein simple; about 48–51 costal crossveins along ScP, all simple; ScP distally fused with RA; subcostal space mostly with uniform width, distally narrow, with about three scp-r crossveins (one or two on its distal portion), proximal one proximad original point of RP); one to five ra-rp crossveins, distal one proximad position of indistinct pterostigma; RP originating near proximal 1/6 of forewing, with eight main branches proximally smooth; six-seven outer gradates, three inner gradates (absence of these developed gradates in left forewing of NIGP171721 might be caused by poor preservation); proximal r-m absent; 1rp-m distad of both original point both of RP and primary branching point of M; M diverging from R proximad of lm-cu; im cell with two-three main branches; 1m-cu distinctly distal to Cu branching point; 2m-cu distinctly distal to cua-cup; CuA nearly pectinately diverged at its distal 1/3, into four main branches; CuP pectinately diverged near its median into 4–5 main branches; A1 pectinately diverged near forewing margin, into 2–4 main branches, each bifurcated or simple; A2 pectinately diverged distally into 3–5 branches; A3 simple or bifurcate.</p><p>Hind wing similar to forewing, but immaculate; costal space markedly narrow, with about 41–46 simple crossveins along ScP; subcostal space slightly narrower, distally with one crossvein; two-three ra-rp crossveins, distal one proximad of position of indistinct pterostigma; original point of RP closer to wing base, with 7–8 branches; two outer gradates, three inner gradates (absence of these developed gradates in hind wings of NIGP171721 might be dissolved during taphonomic processes); 1rp-m sigmoid, distinctly distad original point of RP and proximad primary branching point of M; im cell with two main branches; CuA markedly long, pectinately diverged into 7–9 main branches (almost all distally bifurcated); CuP with long and deep stem, distally short, with 3–4 pectinate simple branches; A1 distally pectinately diverged into 3–4 simple branches; A2 with 3–4 simple, short branches; A3 simple.</p><p>Terminalia bearing dense, long, soft setae. Tergum 9 narrower than ectoprocts, not fused with ectoprocts; sternum 9 absent; gonocoxites 9 paired, large, subtriangular in lateral view, distally simple, rounded and obtuse, bearing long and soft setae, not well-sclerotized but hollowed and partly membranous as some extant berothids; ectoprocts elliptical; callus cerci present and convex.</p><p>Female unknown.</p><p><b>Remarks.</b> <i>Osmyloberotha dispersa</i> <b>sp. nov.</b> can be distinguished from <i>O</i>. <i>chenzuyini</i> <b>sp. nov.</b> by (i) forewing with many dark spots, and (ii) hind wing outer gradates series reduced and incomplete, consisting of only two crossveins, while in <i>O</i>. <i>chenzuyini</i> <b>sp. nov.</b>, (i) the forewing immaculate, and (ii) the hind wing outer gradates are developed and complete, consisting of 10 crossveins. <i>Osmyloberotha dispersa</i> <b>sp. nov.</b> can be distinguished from <i>O</i>. <i>magnimaculata</i> <b>sp. nov.</b> by (i) size of forewing markings subequal, lacking the distinct, large marking, (ii) hind wing immaculate, (iii) hind wing with two series of incomplete gradate crossveins, (iv) fore- and hind wing MP distally with two main branches, while in <i>O</i>. <i>magnimaculata</i> <b>sp. nov.</b>, (i) the forewing has the distinct large marking; (ii) the hind wing is macular, (iii) only with complete outer gradates, and (iv) fore- and hind wing MP distally has three main branches. <i>Osmyloberotha dispersa</i> <b>sp. nov.</b> can be distinguished from <i>O</i>. <i>simpla</i> by (i) forewing markings evenly scattered, (ii) male gonocoxites 9 large, subtriangular in lateral view and (iii) male ectoprocts elliptical, broader than tergum 9, while in <i>O</i>. <i>simpla</i>, there are only 2–4 dark spots arranged along medial longitudinal axis of the forewing; (ii) male gonocoxites 9 are slender, rod-like, and (iii) male ectoprocts are subtriangular and narrower than tergum 9.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_18751352 |
| institution | Zenodo |
| language | |
| publishDate | 2023 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Osmyloberotha dispersa Li, Zhuo, Wang, Nakamine, Yamamoto, Zhang, Ling, Ohl, Aspöck, Aspöck & Liu, 2023, sp. nov. Li, Hong-Yu Zhuo, De Wang, Bo Nakamine, Hiroshi Yamamoto, Shûhei Zhang, Wei-Wei Ling, Jia-Ni Ohl, Michael Aspöck, Ulrike Aspöck, Horst Liu, Xing-Yue Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Berothidae Osmyloberotha Osmyloberotha dispersa <p><b><i>Osmyloberotha dispersa</i> sp. nov.</b></p><p>(Figs 1, 2)</p><p><b>Material.</b> Holotype, ♂, NIGP171721, amber piece polished in the form of a rectangular, transparent cabochon, with length × width about 13.95 × 7.97 mm, height about 4.86 mm (NIGP). Paratype, 1♂, CAM BA-0016, amber piece also preserving a specimen of <i>Jersiberotha</i> sp., three flies, and a beetle; it is polished in the form of a rectangular, transparent cabochon, with length × width about 34.72 × 10.71 mm, height about 5.52 mm (CAM).</p><p><b>Etymology.</b> The specific name “ <i>dispersa</i> ” (Latin, meaning “dispersed”, “scattered”) refers to the evenly dispersed, irregularly shaped, dark markings on the forewing in the new species.</p><p><b>Diagnosis.</b> Forewing with many evenly dispersed, irregular and dark markings. Hind wing immaculate, two incomplete gradates series. Male tergum 9 narrower than ectoprocts; gonocoxites 9 paired, large, subtriangular in lateral view, distally simple and obtuse.</p><p><b>Locality and horizon.</b> Hukawng Valley in Tanai Township, Myitkyina District of Kachin State, Myanmar, Lowermost Cenomanian.</p><p><b>Description.</b> <i>Measurements and proportions.</i> Male. Body length: 2.50–3.34; LP (0.30): WP (0.25) = 1.20; LP: LPT (0.78) = 0.38; LPf (0.45): LPt (0.48) = 0.94; LFW (4.72): WFW (1.60) = 2.95; LHW (4.18): WHW (1.41) = 2.96.</p><p>Male. Head bearing long and soft setae. Vertex moderately domed, bearing markedly long setae. Postocular region markedly narrow in lateral view. Antenna:scape nearly cylindrical,distally slightly swollen, bearing sparse, long setae, 1.5 times as long as wide, and 1.5 times as long as pedicel; pedicel nearly as long as wide, and slightly longer than flagellomere 1; flagellum filiform, with about 33 segments; each flagellomere cylindrical, nearly as long as wide; terminal flagellomere conical, apically tapering into a point. Terminal maxillary and labial palpomeres each proximally distinctly swollen and tapering to an acute point.</p><p>Thorax bearing markedly long setae. Pronotum short, nearly 1/3 length of pterothorax. Forelegs: profemur nearly as long as protibia; protarsus 5-segmented: tarsomere 1 nearly as long as tarsomeres 2–3 combined, tarsomeres 2–3, 5 subequal, tarsomere 4 slightly shorter than tarsomere 3; tarsomeres 1–4 each distally bearing two pairs of thickened setae (spurs). Mid and hind legs similar to forelegs.</p><p>Wings acute, subfalcate, branches of RP mostly distally bifurcate or simple, some trifurcate. Forewing with many evenly dispersed, irregular, dark markings; costal space narrow, not dilated; costal vein distally separate from ScP; ScA absent; recurrent humeral crossvein simple; about 48–51 costal crossveins along ScP, all simple; ScP distally fused with RA; subcostal space mostly with uniform width, distally narrow, with about three scp-r crossveins (one or two on its distal portion), proximal one proximad original point of RP); one to five ra-rp crossveins, distal one proximad position of indistinct pterostigma; RP originating near proximal 1/6 of forewing, with eight main branches proximally smooth; six-seven outer gradates, three inner gradates (absence of these developed gradates in left forewing of NIGP171721 might be caused by poor preservation); proximal r-m absent; 1rp-m distad of both original point both of RP and primary branching point of M; M diverging from R proximad of lm-cu; im cell with two-three main branches; 1m-cu distinctly distal to Cu branching point; 2m-cu distinctly distal to cua-cup; CuA nearly pectinately diverged at its distal 1/3, into four main branches; CuP pectinately diverged near its median into 4–5 main branches; A1 pectinately diverged near forewing margin, into 2–4 main branches, each bifurcated or simple; A2 pectinately diverged distally into 3–5 branches; A3 simple or bifurcate.</p><p>Hind wing similar to forewing, but immaculate; costal space markedly narrow, with about 41–46 simple crossveins along ScP; subcostal space slightly narrower, distally with one crossvein; two-three ra-rp crossveins, distal one proximad of position of indistinct pterostigma; original point of RP closer to wing base, with 7–8 branches; two outer gradates, three inner gradates (absence of these developed gradates in hind wings of NIGP171721 might be dissolved during taphonomic processes); 1rp-m sigmoid, distinctly distad original point of RP and proximad primary branching point of M; im cell with two main branches; CuA markedly long, pectinately diverged into 7–9 main branches (almost all distally bifurcated); CuP with long and deep stem, distally short, with 3–4 pectinate simple branches; A1 distally pectinately diverged into 3–4 simple branches; A2 with 3–4 simple, short branches; A3 simple.</p><p>Terminalia bearing dense, long, soft setae. Tergum 9 narrower than ectoprocts, not fused with ectoprocts; sternum 9 absent; gonocoxites 9 paired, large, subtriangular in lateral view, distally simple, rounded and obtuse, bearing long and soft setae, not well-sclerotized but hollowed and partly membranous as some extant berothids; ectoprocts elliptical; callus cerci present and convex.</p><p>Female unknown.</p><p><b>Remarks.</b> <i>Osmyloberotha dispersa</i> <b>sp. nov.</b> can be distinguished from <i>O</i>. <i>chenzuyini</i> <b>sp. nov.</b> by (i) forewing with many dark spots, and (ii) hind wing outer gradates series reduced and incomplete, consisting of only two crossveins, while in <i>O</i>. <i>chenzuyini</i> <b>sp. nov.</b>, (i) the forewing immaculate, and (ii) the hind wing outer gradates are developed and complete, consisting of 10 crossveins. <i>Osmyloberotha dispersa</i> <b>sp. nov.</b> can be distinguished from <i>O</i>. <i>magnimaculata</i> <b>sp. nov.</b> by (i) size of forewing markings subequal, lacking the distinct, large marking, (ii) hind wing immaculate, (iii) hind wing with two series of incomplete gradate crossveins, (iv) fore- and hind wing MP distally with two main branches, while in <i>O</i>. <i>magnimaculata</i> <b>sp. nov.</b>, (i) the forewing has the distinct large marking; (ii) the hind wing is macular, (iii) only with complete outer gradates, and (iv) fore- and hind wing MP distally has three main branches. <i>Osmyloberotha dispersa</i> <b>sp. nov.</b> can be distinguished from <i>O</i>. <i>simpla</i> by (i) forewing markings evenly scattered, (ii) male gonocoxites 9 large, subtriangular in lateral view and (iii) male ectoprocts elliptical, broader than tergum 9, while in <i>O</i>. <i>simpla</i>, there are only 2–4 dark spots arranged along medial longitudinal axis of the forewing; (ii) male gonocoxites 9 are slender, rod-like, and (iii) male ectoprocts are subtriangular and narrower than tergum 9.</p> |
| title | Osmyloberotha dispersa Li, Zhuo, Wang, Nakamine, Yamamoto, Zhang, Ling, Ohl, Aspöck, Aspöck & Liu, 2023, sp. nov. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Berothidae Osmyloberotha Osmyloberotha dispersa |
| url | https://doi.org/10.5281/zenodo.18751352 |