Kurtodipteromantispa univenula Li, Zhuo, Wang, Nakamine, Yamamoto, Zhang, Ling, Ohl, Aspöck, Aspöck & Liu, 2023, sp. nov.
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2023
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| _version_ | 1866901718519775232 |
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| author | Li, Hong-Yu Zhuo, De Wang, Bo Nakamine, Hiroshi Yamamoto, Shûhei Zhang, Wei-Wei Ling, Jia-Ni Ohl, Michael Aspöck, Ulrike Aspöck, Horst Liu, Xing-Yue |
| author_facet | Li, Hong-Yu Zhuo, De Wang, Bo Nakamine, Hiroshi Yamamoto, Shûhei Zhang, Wei-Wei Ling, Jia-Ni Ohl, Michael Aspöck, Ulrike Aspöck, Horst Liu, Xing-Yue |
| contents | <p><b><i>Kurtodipteromantispa univenula</i> sp. nov.</b></p><p>(Figs 21, 22)</p><p><b>Material.</b> Holotype, ♂, NIGP173316, amber piece also preserving a dipteran; it is polished in the form of an elliptical, transparent cabochon, with length × width about 20.80 × 14.50 mm, height about 3.30 mm (NIGP).</p><p><b>Etymology.</b> The specific name “ <i>univenula</i> ” is a combination of the prefix “ <i>uni</i> -” (Latin, meaning “one”, “single”) and “ <i>venula</i> ” (Latin, meaning “vein”) in reference to the simple forewing MP in the new species, which is distinctive in Dipteromantispidae.</p><p><b>Diagnosis.</b> Profemur stout. Forewing narrow (LFW: WFW = 2.78), immaculate; costal space markedly narrow; 12 simple costal crossveins along ScP; RA branches mostly distally bifurcate; RP with five main branches, only primary one bifurcate; five outer gradates; MA and MP simple; crossveins among CuP, A1–A3 not aligned in a line.</p><p><b>Locality and horizon.</b> Hukawng Valley in Tanai Township, Myitkyina District of Kachin State, Myanmar, Lowermost Cenomanian.</p><p><b>Description.</b> <i>Measurements and proportions</i>. Male. Body length: 5.19; LP (0.59): WP (0.64) = 0.92; LP: LPT (1.42) = 0.42; LPf (1.45): WPf (0.40) = 3.63; LPt (1.03): LPf = 0.71; LFW (4.53): WFW (1.63) = 2.78.</p><p>Male. Head, bearing short and soft setae. Vertex moderately domed; tubercles absent; coronal suture indistinct. Postocular margin narrow in lateral view. Antenna about 0.5–1.0× length of forewing, but shorter than its entire length; scape short and stout, cylindrical, slightly longer than wide, twice as long as pedicel; pedicel short, nearly as long as wide, nearly as long as flagellomere 1; flagellum filiform, with about 58 segments; flagellomeres each cylindrical, nearly as long as wide; terminal flagellomere bullet-like, twice as long as prior one. Terminal maxillary and labial palpomeres each proximally distinctly swollen and tapering to an acute point.</p><p>Thorax bearing short and soft setae. Pronotum short, shield-shaped, posteroventrally not fused, nearly 2/5 length of pterothorax. Profemur stout, proximal distinctly broadened and distally narrow in lateral view, bearing moderately long and soft hair; profemoral ISs tubercle-shaped, only several in posteroventral row visible; major process absent; protibia bearing soft hair shorter than that of profemur, ventrally with about 21 specialized setae, fully developed, continuous and homogeneous, with narrow gaps, flattened and smoothly curved, markedly short; protarsus 5-segmented: ventrally without any specialized setae; tarsomere 1 nearly as long as tarsomeres 2–3 combined; tarsomeres 2–4 subequal; tarsomere 5 nearly as long as tarsomeres 3–4 combined; two pretarsal claws, each simple; arolium sac-like, with rounded apex. Mesothorax enlarged conspicuously, dorsally domed; anterolateral margins of scutum rounded; anterior portion of scutum enlarged and subtriangular, nearly as large as lateral portion, with median suture; mesoscutellum subtriangular, large, slightly shorter than 1/2 length of mesothorax. Metathorax strongly reduced. Mid and hind legs bearing dense long setae; meso- and metatarsi each with tarsomere 1 nearly as long as tarsomeres 2–3 combined, tarsomeres 2–4 subequal, tarsomere 5 slightly shorter tarsomere 4; pretarsal claws and arolium similar to those of foreleg, but larger.</p><p>Forewing narrow, elliptical, immaculate, branches of RP without apical twigging; trichosors absent; costal space markedly narrow; costal vein distally separate from ScP; recurrent humeral crossvein simple; 12 simple costal crossveins along ScP; pterostigma indistinct; ScA absent; ScP distally slightly curved, separate from RA; subcostal space veinless, distinctly broadened distally; RA with nine branches, mostly distally bifurcate; one ra-rp crossvein; RP originating at proximal 1/4 of forewing, strongly angularly curved at ra-rp, with five main branches proximally smooth (only proximal one distally deeply bifurcate, others simple); five outer gradates; 1rp-m distad original point of RP and proximad primary branching point of M; M shortly fused with R distad lm-cu; proximal r-m absent; im cell with two simple branches; MA and MP simple; 2m-cu slightly distal to cua-cup; CuA and CuP simple; A1 deeply bifurcate at middle; A2 and A3 simple; crossveins among CuP, A1-A3 not aligned in a line.</p><p>Hind wing strongly reduced, haltere-like.</p><p>Terminalia bearing long, soft setae. Tergum 9 dorsally broadened, separate from ectoprocts, narrower than tergum 8; sternum 9 developed, shallow, plate-like, slightly longer than sternum 8, not surpassing apex of ectoprocts, about slightly longer than latter; gonocoxites 9 stout, subtriangular, distally simple and obtuse, well-sclerotized; gonocoxites 11 broad, not produced ventrad; ectoprocts subtriangular in dorsal view.</p><p>Female unknown.</p><p><b>Remarks.</b> <i>Kurtodipteromantispa univenula</i> <b>sp. nov.</b> can be distinguished from <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i> by the narrow forewing (LFW: WFW = 2.78) (LFW: WFW = 2.20–2.40 in <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i>), simple MP (deeply bifurcate in <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i>) and male ectoprocts subtriangular in dorsal view (rounded in <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i>).</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_18751391 |
| institution | Zenodo |
| language | |
| publishDate | 2023 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Kurtodipteromantispa univenula Li, Zhuo, Wang, Nakamine, Yamamoto, Zhang, Ling, Ohl, Aspöck, Aspöck & Liu, 2023, sp. nov. Li, Hong-Yu Zhuo, De Wang, Bo Nakamine, Hiroshi Yamamoto, Shûhei Zhang, Wei-Wei Ling, Jia-Ni Ohl, Michael Aspöck, Ulrike Aspöck, Horst Liu, Xing-Yue Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Dipteromantispidae Kurtodipteromantispa Kurtodipteromantispa univenula <p><b><i>Kurtodipteromantispa univenula</i> sp. nov.</b></p><p>(Figs 21, 22)</p><p><b>Material.</b> Holotype, ♂, NIGP173316, amber piece also preserving a dipteran; it is polished in the form of an elliptical, transparent cabochon, with length × width about 20.80 × 14.50 mm, height about 3.30 mm (NIGP).</p><p><b>Etymology.</b> The specific name “ <i>univenula</i> ” is a combination of the prefix “ <i>uni</i> -” (Latin, meaning “one”, “single”) and “ <i>venula</i> ” (Latin, meaning “vein”) in reference to the simple forewing MP in the new species, which is distinctive in Dipteromantispidae.</p><p><b>Diagnosis.</b> Profemur stout. Forewing narrow (LFW: WFW = 2.78), immaculate; costal space markedly narrow; 12 simple costal crossveins along ScP; RA branches mostly distally bifurcate; RP with five main branches, only primary one bifurcate; five outer gradates; MA and MP simple; crossveins among CuP, A1–A3 not aligned in a line.</p><p><b>Locality and horizon.</b> Hukawng Valley in Tanai Township, Myitkyina District of Kachin State, Myanmar, Lowermost Cenomanian.</p><p><b>Description.</b> <i>Measurements and proportions</i>. Male. Body length: 5.19; LP (0.59): WP (0.64) = 0.92; LP: LPT (1.42) = 0.42; LPf (1.45): WPf (0.40) = 3.63; LPt (1.03): LPf = 0.71; LFW (4.53): WFW (1.63) = 2.78.</p><p>Male. Head, bearing short and soft setae. Vertex moderately domed; tubercles absent; coronal suture indistinct. Postocular margin narrow in lateral view. Antenna about 0.5–1.0× length of forewing, but shorter than its entire length; scape short and stout, cylindrical, slightly longer than wide, twice as long as pedicel; pedicel short, nearly as long as wide, nearly as long as flagellomere 1; flagellum filiform, with about 58 segments; flagellomeres each cylindrical, nearly as long as wide; terminal flagellomere bullet-like, twice as long as prior one. Terminal maxillary and labial palpomeres each proximally distinctly swollen and tapering to an acute point.</p><p>Thorax bearing short and soft setae. Pronotum short, shield-shaped, posteroventrally not fused, nearly 2/5 length of pterothorax. Profemur stout, proximal distinctly broadened and distally narrow in lateral view, bearing moderately long and soft hair; profemoral ISs tubercle-shaped, only several in posteroventral row visible; major process absent; protibia bearing soft hair shorter than that of profemur, ventrally with about 21 specialized setae, fully developed, continuous and homogeneous, with narrow gaps, flattened and smoothly curved, markedly short; protarsus 5-segmented: ventrally without any specialized setae; tarsomere 1 nearly as long as tarsomeres 2–3 combined; tarsomeres 2–4 subequal; tarsomere 5 nearly as long as tarsomeres 3–4 combined; two pretarsal claws, each simple; arolium sac-like, with rounded apex. Mesothorax enlarged conspicuously, dorsally domed; anterolateral margins of scutum rounded; anterior portion of scutum enlarged and subtriangular, nearly as large as lateral portion, with median suture; mesoscutellum subtriangular, large, slightly shorter than 1/2 length of mesothorax. Metathorax strongly reduced. Mid and hind legs bearing dense long setae; meso- and metatarsi each with tarsomere 1 nearly as long as tarsomeres 2–3 combined, tarsomeres 2–4 subequal, tarsomere 5 slightly shorter tarsomere 4; pretarsal claws and arolium similar to those of foreleg, but larger.</p><p>Forewing narrow, elliptical, immaculate, branches of RP without apical twigging; trichosors absent; costal space markedly narrow; costal vein distally separate from ScP; recurrent humeral crossvein simple; 12 simple costal crossveins along ScP; pterostigma indistinct; ScA absent; ScP distally slightly curved, separate from RA; subcostal space veinless, distinctly broadened distally; RA with nine branches, mostly distally bifurcate; one ra-rp crossvein; RP originating at proximal 1/4 of forewing, strongly angularly curved at ra-rp, with five main branches proximally smooth (only proximal one distally deeply bifurcate, others simple); five outer gradates; 1rp-m distad original point of RP and proximad primary branching point of M; M shortly fused with R distad lm-cu; proximal r-m absent; im cell with two simple branches; MA and MP simple; 2m-cu slightly distal to cua-cup; CuA and CuP simple; A1 deeply bifurcate at middle; A2 and A3 simple; crossveins among CuP, A1-A3 not aligned in a line.</p><p>Hind wing strongly reduced, haltere-like.</p><p>Terminalia bearing long, soft setae. Tergum 9 dorsally broadened, separate from ectoprocts, narrower than tergum 8; sternum 9 developed, shallow, plate-like, slightly longer than sternum 8, not surpassing apex of ectoprocts, about slightly longer than latter; gonocoxites 9 stout, subtriangular, distally simple and obtuse, well-sclerotized; gonocoxites 11 broad, not produced ventrad; ectoprocts subtriangular in dorsal view.</p><p>Female unknown.</p><p><b>Remarks.</b> <i>Kurtodipteromantispa univenula</i> <b>sp. nov.</b> can be distinguished from <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i> by the narrow forewing (LFW: WFW = 2.78) (LFW: WFW = 2.20–2.40 in <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i>), simple MP (deeply bifurcate in <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i>) and male ectoprocts subtriangular in dorsal view (rounded in <i>K. relicta</i>, <i>K. xiai</i> and <i>K. zhuodei</i>).</p> |
| title | Kurtodipteromantispa univenula Li, Zhuo, Wang, Nakamine, Yamamoto, Zhang, Ling, Ohl, Aspöck, Aspöck & Liu, 2023, sp. nov. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Dipteromantispidae Kurtodipteromantispa Kurtodipteromantispa univenula |
| url | https://doi.org/10.5281/zenodo.18751391 |