| _version_ | 1866901871228092416 |
|---|---|
| author | Pignal, Marc Phillipson, Peter B. Bardot-Vaucoulon, Martine |
| author_facet | Pignal, Marc Phillipson, Peter B. Bardot-Vaucoulon, Martine |
| contents | <p><b>29</b>. <i>Chlorophytum tolyanum</i> Bard.-Vauc. & M.Pignal sp. nov.</p><p>urn:lsid:ipni.org:names:77324054-1</p><p>Figs 5B, 36, 44</p><p><b>Diagnosis</b></p><p>Planta herbacea 60 cm alta, radicibus fasciculatis, unum-tria fusiformia albaque tubercula, (10–15 mm × 2–3 mm), folia grandia (25.5–60 × 3.0– 4.8 cm), duo externa minora, pseudopetiolus 12–17 cm, lamina attenuata ad basim et acuta ad apicem, inferna pagine triste viride (in vivo), margo scaberula, 32 nervis remotis manifestisque. Inflorescentia unica, erecta, ramosa, 42–52 cm alta, pedunculus 19–27 cm longus, bractea inflorescentiae triangularis, 12 × 4 mm, margine leviter scaberula nervisque paulo manifestis. Pedunculus ramificationesque inflorescentiae omnino obtecti pilis brevibus hyalinisque.</p><p>Flores (1–4). Pedicellus articulatus infra medium. Bractea floris triangularis, 3–4 × 2.5 mm, base vaginata acuto apice, 3 nervis paulo manifestis, omnino obtecta pilis brevibus hyalinisque. Perianthium, 5 mm altum. Tepala alba elliptica, subaequalia, 4.9–5.1 × 1 mm, trinervia apice acuminato.</p><p>Stamina 2 coacervati, 4 aequantia perianthum, 2 paulo breviora. Papilla 2/3 filamenti staminum obtegentia (longis staminibus 4.8 mm in longitudine, brevibus, 3.8 mm). Filamentum dorsifixum fere medifixum, antherae lutae, 0.7 × 0.2 mm. Ovarium globosum 0.8 × 0.8 mm. Ovulum 1 per loculum. Stylus filiformis, 6.3 mm.</p><p><b>Etymology</b></p><p>The species is dedicated to Aurélien Toly, ecotourist guide, who was, as ANGAP staff member (today MNP), one of the accompanying guides during the various missions carried out by MBV in the Special Reserve of Ankarana in 1997, 2003 and 2007. He is a co-collector of the material obtained in the field.</p><p><b>Type material</b></p><p>MADAGASCAR – <b>DIANA Region [Antsiranana Prov</b>.<b>]</b> • Ambilobe, RS Ankarana, chemin Amposately au Lac Vert; 12°55′ S, 49°05′ E; 14 Jan. 2003; fl; <i>M. Bardot-Vaucoulon, O. Andrianantoanina, A. Toly & T. Manesy 1193</i>; holotype: P [P00455486]!; isotypes: K!, MO!, TAN!.</p><p><b>Description</b></p><p><i>Plant herbaceous</i>, up to 60 cm tall; <i>roots</i> fasciculate, rigid, bearing 1–3 white fusiform tubers, 10–15 × 2–3 mm; <i>leaves</i> 7, lanceolate, 25.5–60 × 3–4.8 cm, but the 2 outer ones smaller, leaf base attenuate, pseudo-petiole broad, 12–17 cm long, base sheathing, blade dull green below, the margins scabrous, with 28–32, clearly visible, evenly-spaced veins, apices long-tapered and aristate; <i>inflorescence</i> solitary, erect, branched, 42–52 cm in length, peduncle 19–27 cm long; axes entirely covered with short, white, hyaline hairs; subtended by a solitary, triangular bract, 12 × 4 mm, with slightly scabrous margins and barely visible veins; <i>rachis</i> 22–27.5 cm long; <i>flowers</i> 2–4 per node, floral bract 3.5 × 2 mm, triangular, embracing the base, margins scabrous, apex pointed, shorter than the pedicel, veins 3, only faintly visible; pedicel 4.5 mm long, covered with hyaline hairs, the articulation inframedial; <i>perianth</i> 5 mm long, <i>tepals</i> elliptical, white, sub-equal, 4.9–5.1 × ca 1 mm, veins 3, close together; <i>stamens</i> in two unequal groups, 4 long (equaling the perianth), 2 short (slightly smaller), filaments papillose for ca 2/3 of their length, 4.8 mm long for the long ones, 3.8 mm long for the short ones, anthers yellow, 0.7 × 0.2 mm, dorsifixed to submedifixed; <i>gynoecium</i>: ovary 0.8 × 0.8 mm, ovules 1 per locule, style filiform, 6.3 mm long; <i>fruit</i> unknown.</p><p><b>Phenology</b></p><p>The only known collection was in early flower in January, corresponding to the start of the rainy season.</p><p><b>Distribution and ecology</b></p><p>The only known collection of this species is from the Réserve spéciale d’Ankarana [Ankarana Special Reserve] in the North of Madagascar. It occurs in the understory of dry deciduous forest, specifically growing in limestone scree, containing conspicuous large blocks of limestone, with pockets of smaller pieces and humic soil.</p><p><b>Provisional IUCN Red List assessment</b></p><p><i>Chlorophytum tolyanum</i> sp. nov. is known only from a single recent collection in what is regarded as a well-managed Special Reserve, and it is naturally well-protected by its habitat which is difficult to access. However, it is known from only a single occurrence, and it is therefore presumed to be rare. A targeted survey of its occurrence at the site is needed to be able to assess any threats to the species, and in orger to provide a meaningful conservation status, and at present it must be assessed as DD (data deficient).</p><p>A map showing the single occurrence of <i>Chlorophytum tolyanum</i> sp. nov. together with <i>C. ratovosonii</i> sp. nov., another poorly-known species from the north of Madagascar, is provided under the treatment of <i>D. nigrogranulosum</i> sp. nov. (Fig. 36).</p><p><b>Taxonomical remark</b></p><p>This species differs from other species by a mosaic of characters that makes it difficult to understand a differential diagnosis mentioning several other species. We prefer to present a descriptive diagnosis here.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_18762585 |
| institution | Zenodo |
| language | |
| publishDate | 2026 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Chlorophytum tolyanum Bard. - Vauc. & M. Pignal 2026, sp. nov. Pignal, Marc Phillipson, Peter B. Bardot-Vaucoulon, Martine Biodiversity Taxonomy Plantae Tracheophyta Liliopsida Asparagales Asparagaceae Chlorophytum Chlorophytum tolyanum <p><b>29</b>. <i>Chlorophytum tolyanum</i> Bard.-Vauc. & M.Pignal sp. nov.</p><p>urn:lsid:ipni.org:names:77324054-1</p><p>Figs 5B, 36, 44</p><p><b>Diagnosis</b></p><p>Planta herbacea 60 cm alta, radicibus fasciculatis, unum-tria fusiformia albaque tubercula, (10–15 mm × 2–3 mm), folia grandia (25.5–60 × 3.0– 4.8 cm), duo externa minora, pseudopetiolus 12–17 cm, lamina attenuata ad basim et acuta ad apicem, inferna pagine triste viride (in vivo), margo scaberula, 32 nervis remotis manifestisque. Inflorescentia unica, erecta, ramosa, 42–52 cm alta, pedunculus 19–27 cm longus, bractea inflorescentiae triangularis, 12 × 4 mm, margine leviter scaberula nervisque paulo manifestis. Pedunculus ramificationesque inflorescentiae omnino obtecti pilis brevibus hyalinisque.</p><p>Flores (1–4). Pedicellus articulatus infra medium. Bractea floris triangularis, 3–4 × 2.5 mm, base vaginata acuto apice, 3 nervis paulo manifestis, omnino obtecta pilis brevibus hyalinisque. Perianthium, 5 mm altum. Tepala alba elliptica, subaequalia, 4.9–5.1 × 1 mm, trinervia apice acuminato.</p><p>Stamina 2 coacervati, 4 aequantia perianthum, 2 paulo breviora. Papilla 2/3 filamenti staminum obtegentia (longis staminibus 4.8 mm in longitudine, brevibus, 3.8 mm). Filamentum dorsifixum fere medifixum, antherae lutae, 0.7 × 0.2 mm. Ovarium globosum 0.8 × 0.8 mm. Ovulum 1 per loculum. Stylus filiformis, 6.3 mm.</p><p><b>Etymology</b></p><p>The species is dedicated to Aurélien Toly, ecotourist guide, who was, as ANGAP staff member (today MNP), one of the accompanying guides during the various missions carried out by MBV in the Special Reserve of Ankarana in 1997, 2003 and 2007. He is a co-collector of the material obtained in the field.</p><p><b>Type material</b></p><p>MADAGASCAR – <b>DIANA Region [Antsiranana Prov</b>.<b>]</b> • Ambilobe, RS Ankarana, chemin Amposately au Lac Vert; 12°55′ S, 49°05′ E; 14 Jan. 2003; fl; <i>M. Bardot-Vaucoulon, O. Andrianantoanina, A. Toly & T. Manesy 1193</i>; holotype: P [P00455486]!; isotypes: K!, MO!, TAN!.</p><p><b>Description</b></p><p><i>Plant herbaceous</i>, up to 60 cm tall; <i>roots</i> fasciculate, rigid, bearing 1–3 white fusiform tubers, 10–15 × 2–3 mm; <i>leaves</i> 7, lanceolate, 25.5–60 × 3–4.8 cm, but the 2 outer ones smaller, leaf base attenuate, pseudo-petiole broad, 12–17 cm long, base sheathing, blade dull green below, the margins scabrous, with 28–32, clearly visible, evenly-spaced veins, apices long-tapered and aristate; <i>inflorescence</i> solitary, erect, branched, 42–52 cm in length, peduncle 19–27 cm long; axes entirely covered with short, white, hyaline hairs; subtended by a solitary, triangular bract, 12 × 4 mm, with slightly scabrous margins and barely visible veins; <i>rachis</i> 22–27.5 cm long; <i>flowers</i> 2–4 per node, floral bract 3.5 × 2 mm, triangular, embracing the base, margins scabrous, apex pointed, shorter than the pedicel, veins 3, only faintly visible; pedicel 4.5 mm long, covered with hyaline hairs, the articulation inframedial; <i>perianth</i> 5 mm long, <i>tepals</i> elliptical, white, sub-equal, 4.9–5.1 × ca 1 mm, veins 3, close together; <i>stamens</i> in two unequal groups, 4 long (equaling the perianth), 2 short (slightly smaller), filaments papillose for ca 2/3 of their length, 4.8 mm long for the long ones, 3.8 mm long for the short ones, anthers yellow, 0.7 × 0.2 mm, dorsifixed to submedifixed; <i>gynoecium</i>: ovary 0.8 × 0.8 mm, ovules 1 per locule, style filiform, 6.3 mm long; <i>fruit</i> unknown.</p><p><b>Phenology</b></p><p>The only known collection was in early flower in January, corresponding to the start of the rainy season.</p><p><b>Distribution and ecology</b></p><p>The only known collection of this species is from the Réserve spéciale d’Ankarana [Ankarana Special Reserve] in the North of Madagascar. It occurs in the understory of dry deciduous forest, specifically growing in limestone scree, containing conspicuous large blocks of limestone, with pockets of smaller pieces and humic soil.</p><p><b>Provisional IUCN Red List assessment</b></p><p><i>Chlorophytum tolyanum</i> sp. nov. is known only from a single recent collection in what is regarded as a well-managed Special Reserve, and it is naturally well-protected by its habitat which is difficult to access. However, it is known from only a single occurrence, and it is therefore presumed to be rare. A targeted survey of its occurrence at the site is needed to be able to assess any threats to the species, and in orger to provide a meaningful conservation status, and at present it must be assessed as DD (data deficient).</p><p>A map showing the single occurrence of <i>Chlorophytum tolyanum</i> sp. nov. together with <i>C. ratovosonii</i> sp. nov., another poorly-known species from the north of Madagascar, is provided under the treatment of <i>D. nigrogranulosum</i> sp. nov. (Fig. 36).</p><p><b>Taxonomical remark</b></p><p>This species differs from other species by a mosaic of characters that makes it difficult to understand a differential diagnosis mentioning several other species. We prefer to present a descriptive diagnosis here.</p> |
| title | Chlorophytum tolyanum Bard. - Vauc. & M. Pignal 2026, sp. nov. |
| topic | Biodiversity Taxonomy Plantae Tracheophyta Liliopsida Asparagales Asparagaceae Chlorophytum Chlorophytum tolyanum |
| url | https://doi.org/10.5281/zenodo.18762585 |