Diadegma (Nythobia) mirum Klopfstein, 2026, sp. nov.
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2026
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| author | Klopfstein, Seraina |
| author_facet | Klopfstein, Seraina |
| contents | <p><i>Diadegma</i> (<i>Nythobia</i>) <i>mirum</i> sp. nov.</p><p>Fig. 2</p><p><b>Etymology.</b></p><p>The species epithet is derived from the Latin word mīrum, meaning “ wonderful ” – which all of Earth’s creatures ultimately are. The species epithet is an adjective in neuter nominative singular form, as the genus name <i>Diadegma</i> is neuter.</p><p><b>Material examined.</b></p><p><i>Holotype</i>: Switzerland • 1 ♀; Bern, Saanen, Krinnen; 13–14 Sep. 1969; Jacques Aubert leg.; GBIFCH 02040377 (at MZL). <i>Paratype</i>: Sweden • 1 ♀; Kalmar län, Öland, Borgholm, Alvara Dödevi; 57.1872°N, 17.068°E; 12 Jul. 2022; Seraina Klopfstein leg.; NMB - HYMEN 0005569 (at NMB).</p><p><b>Diagnosis.</b></p><p>The new species is one of just five species in the genus in Europe that have the punctation on the lower mesopleuron stronger than the coriaceous background. <i>Diadegma</i> (<i>Nythobia</i>) <i>integrator</i> Aubert, 1970 and <i>D.</i> (<i>Nythobia</i>) <i>rectificator</i> Aubert, 1970 both have tergites 6 and 7 not or weakly excised, while <i>D.</i> (<i>Neoangitia</i>) <i>aculeatum</i> (Bridgman, 1889) has an ovipositor that is much longer than the hind tibia and shiny posterior tergites. The most similar species in Europe with rather strong punctation on the mesopleuron is <i>D.</i> (<i>Nythobia</i>) <i>areolare</i> (Holmgren, 1860), a species known to me only from a single specimen in the Thomson collection in Lund (marked by Jacques F. Aubert as “ TYPE!? ”). The new species can be distinguished from it by its medially orange hind tibia (ivory in <i>D. areolare</i>), the areolet being met by vein 2 m-cu a little apically of its middle (clearly basally in <i>D. areolare</i>), and by the much shorter area superomedia (which is nearly twice longer than wide in <i>D. areolare</i>).</p><p>In Horstmann’s key (Horstmann 1969), this species runs to group III (species with deeply excised tergite 7 and ovipositor sheath shorter than hind tibia), and within the group to either <i>D.</i> (<i>Nythobia</i>) <i>areolare</i>, <i>D.</i> (<i>Nythobia</i>) <i>tenuipes</i> (Thomson, 1887), or <i>D.</i> (<i>Nythobia</i>) <i>combinatum</i> (Holmgren, 1860). While the first of these species is already excluded above, the remaining two have the punctures on the mesopleuron at most as conspicuous as the background sculpture. From <i>D. tenuipes</i>, <i>D. mirum</i> can be further distinguished by its dark fore coxa and trochanter (orange and yellow, respectively, in <i>D. tenuipes</i>) and the higher POL (1.80–1.95 versus 1.25–1.45 in <i>D. tenuipes</i>). The new species shares both these characters with <i>D. combinatum</i> but has its inner eye margins almost parallel ventrally (strongly converging in <i>D. combinatum</i>), a lower OOL (1.05–1.10 versus 1.35–1.50 in <i>D. combinatum</i>), and its ovipositor evenly upcurved (nearly straight in <i>D. combinatum</i>).</p><p>Among the group III species described since 1969, none has a medially orange hind tibia combined with an at least somewhat excised tergite 6 and dark-marked fore trochanters. However, <i>D.</i> (<i>Nythobia</i>) <i>alpinator</i> Aubert, 1970 meets all these criteria and was not included in the 1969 key. It was originally described as a subspecies of <i>D. combinatum</i> (Aubert 1970), later synonymized with that taxon by Horstmann (1973), and only thirty year later resurrected as a separate species (Horstmann 2013). The new species can be distinguished from <i>D. alpinator</i> by the more deeply excised tergite 6, the longer tergite 2 (1.40 versus 1.05–1.15 in <i>D. alpinator</i>), the more distinctly banded hind tibia, and the lower OOL (1.05–1.10 versus 1.25–1.40 in <i>D. alpinator</i>).</p><p><b>Description.</b></p><p>Measurements are based on the two females in the type series, with the value of the holotype repeated in brackets.</p><p>Body 5.1–6.3 (6.3) mm. Temple in lateral view 0.75 times as long as eye, strongly and roundly constricted; OOL 1.05–1.10 (1.05), POL 1.80–1.95 (1.95); face 1.25–1.30 (1.25) times as wide as high, with coriaceous microsculpture and distinct punctures, especially in lower part, matt; malar space 0.5–0.7 (0.7) times width of mandible at base; clypeus a little rounded in profile, with weak microsculpture and distinct punctures; apical margin of clypeus somewhat impressed and reflexed as a weak lamella, margin convex in frontal view. Antenna with 26–28 (28) flagellomeres, about as long as wide or slightly longer in apical third, tapering towards apex.</p><p>Pronotum above punctate on nearly smooth background, in lower half with longitudinal rugae, not bulging below mesoscutum, epomia rather long; mesoscutum coriaceous, with distinct punctures on entire surface; scuto-scutellar groove smooth; mesopleuron only weakly coriaceous, nearly smooth in most parts, with strong and dense punctures, smooth around speculum, anterior to it with longitudinal rugulae amidst the punctation; metapleuron largely smooth and with strong punctures.</p><p>Fore wing 3.25–4.60 (4.60) mm, areolet quadrate, medium-sized to large, strongly petiolate above, with vein 2 m-cu meeting it a bit apically of its middle. Hindwing with nervellus nearly vertical, weakly rounded, unbroken. Hind femur 4.75–4.80 (4.80) times as long as high; hind claw not surpassing pulvillus in lateral view, with strong pecten reaching to more than half its length, a bit shorter than claw itself.</p><p>Propodeum with carinae strong, coriaceous in area basalis and area externa, rugulose in area dentipara and area superomedia, with transverse rugae in area petiolaris; area basalis short, triangular, directly meeting area superomedia; area superomedia of average size, long pentagonal with lateral sides slightly converging, a bit longer than wide, posteriorly open; area petiolaris very weakly impressed medially.</p><p>Metasoma laterally compressed, widest around tergite 2 or 3, coriaceous and with weak but dense punctures. Tergite 1 with small, deep glymma a bit closer to spiracle than to base, postpetiolus first rounded, then flat dorsally. Tergite 2 1.40–1.45 (1.40) times longer than wide at apex, tergites 6 and 7 deeply excised. Ovipositor sheath 0.85–0.90 (0.90) times as long as hind tibia, evenly upcurved, with notch rather distant from apex, ventral valve apparently smooth.</p><p>Black; palps, mandibles except teeth, tegula, wing base, and fore trochantellus yellow; scape black; wing venation brown, pterostigma medium brown. All coxae black; trochanters and mid and hind trochantelli black at base, narrowly yellow at apex; femora light to dark orange, ventrally marked with black at least at base; fore and mid tibiae orange, mid tibia with indication of subbasal and apical brown marks, tarsi orange to brown; hind tibia on the inner side orange, on the outer ivory at base, orange medially, dark brown to black subbasally and apically; hind spurs and base of first tarsomere orange to brown, remainder dark brown. Metasoma entirely black.</p><p>No host data is available for this species.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_19092378 |
| institution | Zenodo |
| language | |
| publishDate | 2026 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Diadegma (Nythobia) mirum Klopfstein, 2026, sp. nov. Klopfstein, Seraina Biodiversity Taxonomy Animalia Arthropoda Insecta Hymenoptera Ichneumonidae Diadegma Diadegma mirum <p><i>Diadegma</i> (<i>Nythobia</i>) <i>mirum</i> sp. nov.</p><p>Fig. 2</p><p><b>Etymology.</b></p><p>The species epithet is derived from the Latin word mīrum, meaning “ wonderful ” – which all of Earth’s creatures ultimately are. The species epithet is an adjective in neuter nominative singular form, as the genus name <i>Diadegma</i> is neuter.</p><p><b>Material examined.</b></p><p><i>Holotype</i>: Switzerland • 1 ♀; Bern, Saanen, Krinnen; 13–14 Sep. 1969; Jacques Aubert leg.; GBIFCH 02040377 (at MZL). <i>Paratype</i>: Sweden • 1 ♀; Kalmar län, Öland, Borgholm, Alvara Dödevi; 57.1872°N, 17.068°E; 12 Jul. 2022; Seraina Klopfstein leg.; NMB - HYMEN 0005569 (at NMB).</p><p><b>Diagnosis.</b></p><p>The new species is one of just five species in the genus in Europe that have the punctation on the lower mesopleuron stronger than the coriaceous background. <i>Diadegma</i> (<i>Nythobia</i>) <i>integrator</i> Aubert, 1970 and <i>D.</i> (<i>Nythobia</i>) <i>rectificator</i> Aubert, 1970 both have tergites 6 and 7 not or weakly excised, while <i>D.</i> (<i>Neoangitia</i>) <i>aculeatum</i> (Bridgman, 1889) has an ovipositor that is much longer than the hind tibia and shiny posterior tergites. The most similar species in Europe with rather strong punctation on the mesopleuron is <i>D.</i> (<i>Nythobia</i>) <i>areolare</i> (Holmgren, 1860), a species known to me only from a single specimen in the Thomson collection in Lund (marked by Jacques F. Aubert as “ TYPE!? ”). The new species can be distinguished from it by its medially orange hind tibia (ivory in <i>D. areolare</i>), the areolet being met by vein 2 m-cu a little apically of its middle (clearly basally in <i>D. areolare</i>), and by the much shorter area superomedia (which is nearly twice longer than wide in <i>D. areolare</i>).</p><p>In Horstmann’s key (Horstmann 1969), this species runs to group III (species with deeply excised tergite 7 and ovipositor sheath shorter than hind tibia), and within the group to either <i>D.</i> (<i>Nythobia</i>) <i>areolare</i>, <i>D.</i> (<i>Nythobia</i>) <i>tenuipes</i> (Thomson, 1887), or <i>D.</i> (<i>Nythobia</i>) <i>combinatum</i> (Holmgren, 1860). While the first of these species is already excluded above, the remaining two have the punctures on the mesopleuron at most as conspicuous as the background sculpture. From <i>D. tenuipes</i>, <i>D. mirum</i> can be further distinguished by its dark fore coxa and trochanter (orange and yellow, respectively, in <i>D. tenuipes</i>) and the higher POL (1.80–1.95 versus 1.25–1.45 in <i>D. tenuipes</i>). The new species shares both these characters with <i>D. combinatum</i> but has its inner eye margins almost parallel ventrally (strongly converging in <i>D. combinatum</i>), a lower OOL (1.05–1.10 versus 1.35–1.50 in <i>D. combinatum</i>), and its ovipositor evenly upcurved (nearly straight in <i>D. combinatum</i>).</p><p>Among the group III species described since 1969, none has a medially orange hind tibia combined with an at least somewhat excised tergite 6 and dark-marked fore trochanters. However, <i>D.</i> (<i>Nythobia</i>) <i>alpinator</i> Aubert, 1970 meets all these criteria and was not included in the 1969 key. It was originally described as a subspecies of <i>D. combinatum</i> (Aubert 1970), later synonymized with that taxon by Horstmann (1973), and only thirty year later resurrected as a separate species (Horstmann 2013). The new species can be distinguished from <i>D. alpinator</i> by the more deeply excised tergite 6, the longer tergite 2 (1.40 versus 1.05–1.15 in <i>D. alpinator</i>), the more distinctly banded hind tibia, and the lower OOL (1.05–1.10 versus 1.25–1.40 in <i>D. alpinator</i>).</p><p><b>Description.</b></p><p>Measurements are based on the two females in the type series, with the value of the holotype repeated in brackets.</p><p>Body 5.1–6.3 (6.3) mm. Temple in lateral view 0.75 times as long as eye, strongly and roundly constricted; OOL 1.05–1.10 (1.05), POL 1.80–1.95 (1.95); face 1.25–1.30 (1.25) times as wide as high, with coriaceous microsculpture and distinct punctures, especially in lower part, matt; malar space 0.5–0.7 (0.7) times width of mandible at base; clypeus a little rounded in profile, with weak microsculpture and distinct punctures; apical margin of clypeus somewhat impressed and reflexed as a weak lamella, margin convex in frontal view. Antenna with 26–28 (28) flagellomeres, about as long as wide or slightly longer in apical third, tapering towards apex.</p><p>Pronotum above punctate on nearly smooth background, in lower half with longitudinal rugae, not bulging below mesoscutum, epomia rather long; mesoscutum coriaceous, with distinct punctures on entire surface; scuto-scutellar groove smooth; mesopleuron only weakly coriaceous, nearly smooth in most parts, with strong and dense punctures, smooth around speculum, anterior to it with longitudinal rugulae amidst the punctation; metapleuron largely smooth and with strong punctures.</p><p>Fore wing 3.25–4.60 (4.60) mm, areolet quadrate, medium-sized to large, strongly petiolate above, with vein 2 m-cu meeting it a bit apically of its middle. Hindwing with nervellus nearly vertical, weakly rounded, unbroken. Hind femur 4.75–4.80 (4.80) times as long as high; hind claw not surpassing pulvillus in lateral view, with strong pecten reaching to more than half its length, a bit shorter than claw itself.</p><p>Propodeum with carinae strong, coriaceous in area basalis and area externa, rugulose in area dentipara and area superomedia, with transverse rugae in area petiolaris; area basalis short, triangular, directly meeting area superomedia; area superomedia of average size, long pentagonal with lateral sides slightly converging, a bit longer than wide, posteriorly open; area petiolaris very weakly impressed medially.</p><p>Metasoma laterally compressed, widest around tergite 2 or 3, coriaceous and with weak but dense punctures. Tergite 1 with small, deep glymma a bit closer to spiracle than to base, postpetiolus first rounded, then flat dorsally. Tergite 2 1.40–1.45 (1.40) times longer than wide at apex, tergites 6 and 7 deeply excised. Ovipositor sheath 0.85–0.90 (0.90) times as long as hind tibia, evenly upcurved, with notch rather distant from apex, ventral valve apparently smooth.</p><p>Black; palps, mandibles except teeth, tegula, wing base, and fore trochantellus yellow; scape black; wing venation brown, pterostigma medium brown. All coxae black; trochanters and mid and hind trochantelli black at base, narrowly yellow at apex; femora light to dark orange, ventrally marked with black at least at base; fore and mid tibiae orange, mid tibia with indication of subbasal and apical brown marks, tarsi orange to brown; hind tibia on the inner side orange, on the outer ivory at base, orange medially, dark brown to black subbasally and apically; hind spurs and base of first tarsomere orange to brown, remainder dark brown. Metasoma entirely black.</p><p>No host data is available for this species.</p> |
| title | Diadegma (Nythobia) mirum Klopfstein, 2026, sp. nov. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Hymenoptera Ichneumonidae Diadegma Diadegma mirum |
| url | https://doi.org/10.5281/zenodo.19092378 |