Athysanini Van Duzee 1892
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2026
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| _version_ | 1866901607347650560 |
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| author | Zahniser, James N. Dietrich, Christopher H. Cao, Yanghui Dmitriev, Dmitry A. Pinedo-Escatel, J. Adilson |
| author_facet | Zahniser, James N. Dietrich, Christopher H. Cao, Yanghui Dmitriev, Dmitry A. Pinedo-Escatel, J. Adilson |
| contents | <p><b>Tribe Athysanini Van Duzee, 1892</b></p><p><b>Remarks</b></p><p>This tribe has long been treated as a taxonomic dumping ground for genera of Deltocephalinae that do not fit the definitions of the smaller, morphologically better-defined tribes included in the subfamily. Previous phylogenetic analyses based on both morphology (Zahniser & Dietrich 2008) and molecular data (Zahniser & Dietrich 2010, 2013; Cao <i>et al</i>. 2022) have shown that the tribe, as defined by previous authors, e.g., Oman <i>et al</i>. (1990) and Zahniser & Dietrich (2013), is polyphyletic. The phylogenomic results of Cao <i>et al</i>. (2022) (Fig. 1; Supp. file 2) recovered several independent lineages of genera previously included in Athysanini. Among the larger of such groups is a clade that includes the endemic New World tribes Bahitini, Pendarini and Scaphytopiini as well as most of the endemic Neotropical genera previously placed in Athysanini. The latter genera are here included in the revalidated tribe Platymetopiini (see below). Another large clade includes the mostly Indomalayan bamboo-specialist tribes Mukariini and Vartini. These two clades and their included tribes and subtribes are treated separately below.</p><p>A third large clade recovered by Cao <i>et al</i>. (2022) comprises Athysanina as recognized here (= the <i>Athysanus</i> group sensu Zahniser & Dietrich 2013) as well as two groups which are newly considered subtribes of Athysanini here, the western Nearctic subtribe Cochlorhinina and the Holarctic subtribe Koebeliina. Cochlorhinina shares with some Athysanina one or two dorsal teeth on the fused portion of the second valvulae, and some (e.g., <i>Gloridonus</i> Ball, 1936) share a relatively broad frontoclypeus and parallel-sided anteclypeus with Athysanina. Some members of Koebeliina are relatively derived morphologically, possessing an elongated and flattened head which is perhaps related to their unusual habit of specialization on pines; they do not share obvious characters with Athysanina other than the presumably symplesiomorphic Y-shaped connective articulated to the aedeagus. Koebeliina and some Cochlorhinina share the presence of platellae on the plantar surface of metatarsomere I and some Koebeliina, e.g., <i>Grypotes puncticollis</i> (Herrich-Schäffer, 1834), share with Cochlorhinina a Y-shaped connective fused to the aedeagus. The characterizations of Cochlorhinina and Koebeliina remain unchanged from their tribal diagnoses and descriptions (Dietrich & Dmitriev 2003; Zahniser & Dietrich 2013). Grypotinae Haupt, 1929 syn. nov., a small group of five genera, was previously included as a subtribe of Koebeliini. It is here considered a synonym of Koebeliina, previously containing only the type genus, with which it shares several morphological synapomorphies (Dietrich & Dmitriev 2003) and host plant specialization on <i>Pinus</i> spp. All recent phylogenetic analyses of Deltocephalinae strongly support a close relationship between <i>Koebelia</i> Baker, 1897 and sampled genera of Grypotinae. Cao <i>et al</i>. (2022) recovered one clade of Athysanina (including <i>Athysanus</i> Burmeister, 1838) as sister to Cochlorhinina and another (including <i>Anoterostemma</i> Löw, 1885, <i>Thamnotettix</i> Zetterstedt, 1837 and <i>Euscelis</i> Brullé, 1832) as sister to Koebeliina. Although it may eventually be possible to recognize both clades of Athysanina as separate subtribes, consistent morphological characters that distinguish members of these two clades have not been found; thus, they are here considered to belong to a single subtribe, Athysanina.</p><p>Tribe Athysanini, as here recognized, although well-supported as a monophyletic group, remains somewhat morphologically heterogeneous and difficult to diagnose based on a few morphological characters. Thus, we have not attempted a formal re-diagnosis of the tribe, although most included genera may be recognized by the characters provided for the nominotypical subtribe below. As mentioned above, subtribes Cochlorhinina and Koebeliina are smaller groups that share the Y-shaped connective found in Athysanina but each have distinctive characters that easily separate them from other Athysanini.</p><p><b>Distribution</b></p><p>Nearctic, Palearctic, Oriental.</p><p><b>Included subtribes</b></p><p>Athysanina, Cochlorhinina stat. nov., Koebeliina stat. nov. (= Grypotinae Haupt syn. nov.).</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_19240008 |
| institution | Zenodo |
| language | |
| publishDate | 2026 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Athysanini Van Duzee 1892 Zahniser, James N. Dietrich, Christopher H. Cao, Yanghui Dmitriev, Dmitry A. Pinedo-Escatel, J. Adilson Biodiversity Taxonomy Animalia Arthropoda Insecta Hemiptera Cicadellidae <p><b>Tribe Athysanini Van Duzee, 1892</b></p><p><b>Remarks</b></p><p>This tribe has long been treated as a taxonomic dumping ground for genera of Deltocephalinae that do not fit the definitions of the smaller, morphologically better-defined tribes included in the subfamily. Previous phylogenetic analyses based on both morphology (Zahniser & Dietrich 2008) and molecular data (Zahniser & Dietrich 2010, 2013; Cao <i>et al</i>. 2022) have shown that the tribe, as defined by previous authors, e.g., Oman <i>et al</i>. (1990) and Zahniser & Dietrich (2013), is polyphyletic. The phylogenomic results of Cao <i>et al</i>. (2022) (Fig. 1; Supp. file 2) recovered several independent lineages of genera previously included in Athysanini. Among the larger of such groups is a clade that includes the endemic New World tribes Bahitini, Pendarini and Scaphytopiini as well as most of the endemic Neotropical genera previously placed in Athysanini. The latter genera are here included in the revalidated tribe Platymetopiini (see below). Another large clade includes the mostly Indomalayan bamboo-specialist tribes Mukariini and Vartini. These two clades and their included tribes and subtribes are treated separately below.</p><p>A third large clade recovered by Cao <i>et al</i>. (2022) comprises Athysanina as recognized here (= the <i>Athysanus</i> group sensu Zahniser & Dietrich 2013) as well as two groups which are newly considered subtribes of Athysanini here, the western Nearctic subtribe Cochlorhinina and the Holarctic subtribe Koebeliina. Cochlorhinina shares with some Athysanina one or two dorsal teeth on the fused portion of the second valvulae, and some (e.g., <i>Gloridonus</i> Ball, 1936) share a relatively broad frontoclypeus and parallel-sided anteclypeus with Athysanina. Some members of Koebeliina are relatively derived morphologically, possessing an elongated and flattened head which is perhaps related to their unusual habit of specialization on pines; they do not share obvious characters with Athysanina other than the presumably symplesiomorphic Y-shaped connective articulated to the aedeagus. Koebeliina and some Cochlorhinina share the presence of platellae on the plantar surface of metatarsomere I and some Koebeliina, e.g., <i>Grypotes puncticollis</i> (Herrich-Schäffer, 1834), share with Cochlorhinina a Y-shaped connective fused to the aedeagus. The characterizations of Cochlorhinina and Koebeliina remain unchanged from their tribal diagnoses and descriptions (Dietrich & Dmitriev 2003; Zahniser & Dietrich 2013). Grypotinae Haupt, 1929 syn. nov., a small group of five genera, was previously included as a subtribe of Koebeliini. It is here considered a synonym of Koebeliina, previously containing only the type genus, with which it shares several morphological synapomorphies (Dietrich & Dmitriev 2003) and host plant specialization on <i>Pinus</i> spp. All recent phylogenetic analyses of Deltocephalinae strongly support a close relationship between <i>Koebelia</i> Baker, 1897 and sampled genera of Grypotinae. Cao <i>et al</i>. (2022) recovered one clade of Athysanina (including <i>Athysanus</i> Burmeister, 1838) as sister to Cochlorhinina and another (including <i>Anoterostemma</i> Löw, 1885, <i>Thamnotettix</i> Zetterstedt, 1837 and <i>Euscelis</i> Brullé, 1832) as sister to Koebeliina. Although it may eventually be possible to recognize both clades of Athysanina as separate subtribes, consistent morphological characters that distinguish members of these two clades have not been found; thus, they are here considered to belong to a single subtribe, Athysanina.</p><p>Tribe Athysanini, as here recognized, although well-supported as a monophyletic group, remains somewhat morphologically heterogeneous and difficult to diagnose based on a few morphological characters. Thus, we have not attempted a formal re-diagnosis of the tribe, although most included genera may be recognized by the characters provided for the nominotypical subtribe below. As mentioned above, subtribes Cochlorhinina and Koebeliina are smaller groups that share the Y-shaped connective found in Athysanina but each have distinctive characters that easily separate them from other Athysanini.</p><p><b>Distribution</b></p><p>Nearctic, Palearctic, Oriental.</p><p><b>Included subtribes</b></p><p>Athysanina, Cochlorhinina stat. nov., Koebeliina stat. nov. (= Grypotinae Haupt syn. nov.).</p> |
| title | Athysanini Van Duzee 1892 |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Hemiptera Cicadellidae |
| url | https://doi.org/10.5281/zenodo.19240008 |