Hellerium Cadena-Castañeda, 2026, gen. nov.
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| author | Cadena-Castañeda, Oscar J. |
| author_facet | Cadena-Castañeda, Oscar J. |
| contents | <p><b><i>Hellerium</i> gen. nov.</b></p><p><b>Taxa included.</b> <i>Hellerium schoenemanni</i> (Karsch, 1889) <b>comb. nov.</b></p><p><b>Description.</b> Body slender and small (10–11 mm) (Fig. 17G). Coloration pale green or yellowish; pronotal disc bordered laterally by a distinct white stripe; thorax and abdomen covered with numerous small reddish punctuations. <b>Head</b> rounded; fastigium of the vertex slightly narrower than the first antennal segment in both frontal and dorsal views; eyes circular and prominent. <b>Thorax.</b> Pronotum lacking lateral carinae and humeral notch (Fig. 18K); lateral margins of the pronotal disc parallel, without median constrictions; anterior and posterior margins nearly straight (Fig. 18L). Lateral lobes wider than tall, with uniformly rounded posterior and ventral margins (Fig. 18K). Auditory spiracle of moderate size, partially exposed at the ventral edge of the pronotal lateral lobe (Fig. 18K). <b>Wings.</b> Tegmina short, not extending beyond the third or fourth abdominal segment (Fig. 18L); hind wings absent. <b>Legs.</b> Fore coxae unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora distinctly surpass the apex of the abdomen. <b>Abdomen.</b> Last abdominal tergite strongly inflated and ventrally deflexed, almost entirely concealing the cerci (Fig. 19L). Cerci with a long, medially directed preapical spine (Fig. 19M). Subgenital plate rectangular and spatulate, with a broad V-shaped notch at the apex; styli absent and without conical, styliform projections.</p><p><b>Female.</b> Similar in general appearance and size to the male (Fig. 19H). Tegmina ovoid, overlapping dorsally, not surpassing the second abdominal segment. Cerci conical, robust, and short; epiproct triangular, nearly as wide as long, apex rounded. Ovipositor more than twice the length of the pronotal disc, nearly straight, slightly curved upward at the distal half, with finely denticulated margins. The basal ridge of the ventral valve of the ovipositor is triangular and modified (Fig. 19H). Subgenital plate triangular, as wide as long, with a truncate apex.</p><p><b>Species included.</b> Only the type species.</p><p><b>Distribution.</b> Central Chile (Map 7).</p><p><b>Etymology.</b> The genus is dedicated to the orthopterist Klaus-Gerhard Heller, in recognition of his valuable contributions, particularly to the study of phaneropterines in Europe and Africa, as well as his significant work in orthopteran bioacoustics. The gender of this name is being established as neuter.</p><p><b>Comparison.</b> <i>Hellerium</i> <b>gen. nov.</b> differs from <i>Anisophya</i> (<i>Castellium</i>) <b>subgen. nov.</b> in the females’ long ovipositor, a character that aligns it more closely with <i>Anisophya</i> s.s. species. However, <i>Hellerium</i> <b>gen. nov.</b> is distinguished from <i>Anisophya</i> by its fastigium, which is narrower than the antennal scape. Males of <i>Hellerium</i> <b>gen. nov.</b> are easily separated from those of <i>Anisophya</i> s.s. and <i>Anisophya</i> (<i>Castellium</i>) <b>subgen. nov.</b> by their terminalia: in the latter two subgenera, the cerci lack an elongated preapical spine, and the tenth tergite does not cover most of the cerci.</p><p>Females of <i>Hellerium</i> <b>gen. nov.</b> also differ in a triangular and modified basal ridge on the ventral valve of the ovipositor, this condition is not observed in the other compared genera, where the basal ridge is represented only by a narrow, normally developed fold.</p><p><b>Comments.</b> The new genus includes <i>H. schoenemanni</i> <b>comb. nov.</b>, one of the three Chilean species previously placed in <i>Anisophya</i>, thereby excluding the presence of that genus in Chile. As previously discussed, <i>Hellerium</i> <b>gen. nov.</b> clearly differs from <i>Anisophya</i>. This taxon appears to be the only member of the tribe that has managed to cross to the western side of the Andes, a region where species of the tribes Cosmophyllini and Aniarellini <b>trib. nov.</b>, considered by some authors to be the most “primitive” in South America, are more commonly found (Nickle 2011).</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_19559804 |
| institution | Zenodo |
| language | |
| publishDate | 2026 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Hellerium Cadena-Castañeda, 2026, gen. nov. Cadena-Castañeda, Oscar J. Biodiversity Taxonomy Animalia Arthropoda Insecta Orthoptera Tettigoniidae Hellerium <p><b><i>Hellerium</i> gen. nov.</b></p><p><b>Taxa included.</b> <i>Hellerium schoenemanni</i> (Karsch, 1889) <b>comb. nov.</b></p><p><b>Description.</b> Body slender and small (10–11 mm) (Fig. 17G). Coloration pale green or yellowish; pronotal disc bordered laterally by a distinct white stripe; thorax and abdomen covered with numerous small reddish punctuations. <b>Head</b> rounded; fastigium of the vertex slightly narrower than the first antennal segment in both frontal and dorsal views; eyes circular and prominent. <b>Thorax.</b> Pronotum lacking lateral carinae and humeral notch (Fig. 18K); lateral margins of the pronotal disc parallel, without median constrictions; anterior and posterior margins nearly straight (Fig. 18L). Lateral lobes wider than tall, with uniformly rounded posterior and ventral margins (Fig. 18K). Auditory spiracle of moderate size, partially exposed at the ventral edge of the pronotal lateral lobe (Fig. 18K). <b>Wings.</b> Tegmina short, not extending beyond the third or fourth abdominal segment (Fig. 18L); hind wings absent. <b>Legs.</b> Fore coxae unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora distinctly surpass the apex of the abdomen. <b>Abdomen.</b> Last abdominal tergite strongly inflated and ventrally deflexed, almost entirely concealing the cerci (Fig. 19L). Cerci with a long, medially directed preapical spine (Fig. 19M). Subgenital plate rectangular and spatulate, with a broad V-shaped notch at the apex; styli absent and without conical, styliform projections.</p><p><b>Female.</b> Similar in general appearance and size to the male (Fig. 19H). Tegmina ovoid, overlapping dorsally, not surpassing the second abdominal segment. Cerci conical, robust, and short; epiproct triangular, nearly as wide as long, apex rounded. Ovipositor more than twice the length of the pronotal disc, nearly straight, slightly curved upward at the distal half, with finely denticulated margins. The basal ridge of the ventral valve of the ovipositor is triangular and modified (Fig. 19H). Subgenital plate triangular, as wide as long, with a truncate apex.</p><p><b>Species included.</b> Only the type species.</p><p><b>Distribution.</b> Central Chile (Map 7).</p><p><b>Etymology.</b> The genus is dedicated to the orthopterist Klaus-Gerhard Heller, in recognition of his valuable contributions, particularly to the study of phaneropterines in Europe and Africa, as well as his significant work in orthopteran bioacoustics. The gender of this name is being established as neuter.</p><p><b>Comparison.</b> <i>Hellerium</i> <b>gen. nov.</b> differs from <i>Anisophya</i> (<i>Castellium</i>) <b>subgen. nov.</b> in the females’ long ovipositor, a character that aligns it more closely with <i>Anisophya</i> s.s. species. However, <i>Hellerium</i> <b>gen. nov.</b> is distinguished from <i>Anisophya</i> by its fastigium, which is narrower than the antennal scape. Males of <i>Hellerium</i> <b>gen. nov.</b> are easily separated from those of <i>Anisophya</i> s.s. and <i>Anisophya</i> (<i>Castellium</i>) <b>subgen. nov.</b> by their terminalia: in the latter two subgenera, the cerci lack an elongated preapical spine, and the tenth tergite does not cover most of the cerci.</p><p>Females of <i>Hellerium</i> <b>gen. nov.</b> also differ in a triangular and modified basal ridge on the ventral valve of the ovipositor, this condition is not observed in the other compared genera, where the basal ridge is represented only by a narrow, normally developed fold.</p><p><b>Comments.</b> The new genus includes <i>H. schoenemanni</i> <b>comb. nov.</b>, one of the three Chilean species previously placed in <i>Anisophya</i>, thereby excluding the presence of that genus in Chile. As previously discussed, <i>Hellerium</i> <b>gen. nov.</b> clearly differs from <i>Anisophya</i>. This taxon appears to be the only member of the tribe that has managed to cross to the western side of the Andes, a region where species of the tribes Cosmophyllini and Aniarellini <b>trib. nov.</b>, considered by some authors to be the most “primitive” in South America, are more commonly found (Nickle 2011).</p> |
| title | Hellerium Cadena-Castañeda, 2026, gen. nov. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Orthoptera Tettigoniidae Hellerium |
| url | https://doi.org/10.5281/zenodo.19559804 |