Sierola allithoros Magnacca, 2026, sp. nov.
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| author | Magnacca, Karl |
| author_facet | Magnacca, Karl |
| contents | <p><b><i>Sierola allithoros</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: E30FF356-4328-493B-9D9C-4BE72F898135</p><p>Figs 10, 15</p><p><b>Diagnosis</b></p><p>Belonging to the complex of species with the clypeus moderately large and quadrate, visibly parallel-sided beyond the antennae. Forming a species pair with <i>S. campanula</i> sp. nov., united in having the frons very narrow (WF/HE ~0.8), clypeus with a low but distinct medial carina and broadly rounded apically rather than straight truncate or emarginate, vein Rs ending short of the anterior wing margin, and S5 with a pair of elongate setae. Separated by having the mandible reddish brown and slightly twisted, propodeum with the median carina faint, and profemur mostly yellow.</p><p><b>Etymology</b></p><p>From the Greek ‘ <i>allithoros</i> ’, ‘cross-eyed’ or ‘squinting’, from the dorsally close eyes. It is a noun in apposition.</p><p><b>Type material</b></p><p><b>Holotype</b></p><p>AUSTRALIA – <b>Queensland</b> • ♀; Upper Clayton Gully, near Cunningham’s Gap; elev. 400–600 m; 25 Apr. 1974; I. Naumann leg.; sweeping long grass; wet sclerophyll forest; QM T262151.</p><p><b>Paratypes</b></p><p>AUSTRALIA – <b>New South Wales</b> • 2 ♀♀, 1 ♂; Hat Head; 31.0626° S, 153.052° E; elev. 37 m; 4–18 Feb. 2018; P. Hebert leg.; Malaise trap, trap #3; BOLD: NSWHO3417-18, NSWHO3816-18, NSWHO3706-18, BIN ADV9054; ANIC. – <b>Queensland</b> • 1 ♀; Corroy [sic: Cooroy]; 15 Sep. 1979; G. Gordh leg.; UCRC 401812 • 1 ♀; Acacia Ridge near Brisbane; 13 Dec. 1979; G. Gordh leg.; UCRC 556776.</p><p><b>Description</b></p><p><b>Female</b></p><p>HEAD. Clypeal process semicircular, apex broadly rounded; dorsally with a distinct but narrow, sharp ridge raised slightly above the otherwise flat surface; in profile evenly sloping, nearly straight. Mandible moderately broad, more or less parallel-sided except at base; slightly twisted, outwardly shining but with distinct, fine microsculpture; in ventral view straight or nearly so. Apex of mandible slightly oblique, with teeth distinctly stronger ventrally. Malar space very small. Dorsal setae of head shorter than flagellar width. Frons with moderately close punctures between antennae and midpoint of eyes, 1–3 pit widths apart, sparse posteriorly. Vertex and gena setae similar to and only slightly longer than setae of frons. Frons microsculpture finely microreticulate to very weak and mostly smooth, shining. Vertex in dorsal view more or less straight; occipital margin broadly rounded. Lateral ocelli distinctly separate from vertex crest by less than ocellus length. Ocellar triangle anterior angle strongly acute. Head more or less parallel-sided for some distance posterior to eyes. Eye setae short, only about as long as 1–2 ommatidia. Dorsum of head in lateral view slightly convex, head deepest on line through posterior margin of eye. Venter of head posterior to mandible convex in lateral view, with relatively sparse setae. Head weakly convex ventrally, genal angle absent. Side of head posterior to eye with a line of smooth tesselate microsculpture extending from posteroventral margin of eye to vertex crest, distinctly different from smoother sculpting dorsally and ventrally. Gena ventral to eye convex; sculpture coriaceous ventrally and posterolaterally, with broad band of tesselate-colliculate sculpture extending diagonally on each side from anterior margin of eye to posteroventral margin of head. Postgenal depression absent. Antenna extending to anterior margin of pronotum.</p><p>MESOSOMA. Anterior margin of dorsal pronotal area in lateral view weakly angulate. Pronotal setae about as long as flagellar width or scarcely longer. Propodeal disc laterally coarsely rugulose; median carina faintly present; transverse carina absent, brow of declivity broadly rounded; metapleural carinae complete. Profemur strongly broadened, less than twice as long as wide, strongly laterally compressed, slightly arcuate in dorsal view.</p><p>METASOMA. Shining, apical tergites punctate laterally. S5 with a pair of long erect setae. Sixth sternum broadly glabrous medially, with sparse setae laterally.</p><p>COLOR. Body entirely dark brown to black. Antenna with scape and base of flagellum pale, becoming darker over apical half. Mandible reddish brown. Procoxa and femur yellow except dorsal margin of femur, meso- and metafemora brown, tibiae, tarsi, and trochanters yellow.</p><p>WINGS. Membrane hyaline. Basal cells of fore wing nearly glabrous, with only a few scattered setae. Cell 1M weakly convex posteriorly, corner posteroapical. Distal portion of vein Rs nearly straight but not quite meeting anterior wing margin, poststigmal abscissa of R1 indistinct.</p><p>MEASUREMENTS. LFW 1.15 mm; LM 0.62 mm (0.59–0.69); WH/LH 0.74 (0.70–0.74); DH/LH 0.52 (0.50–0.52); EV/HE 0.70 (0.64–0.76); EV/WF 0.88 (0.84–1.00); WF/HE 0.79 (0.76–0.84); WH/WF 2.18 (2.05–2.24); OOL/WOT 1.71 (1.67–1.88); DM/LM 0.40; FL/FW 1.96 (1.96–2.08); HFL/HFW 2.16 (2.00–2.33); A3 L/ W 0.83 (0.83–1.00); 2R1 L/ W 3.14. N = 5.</p><p><b>Male</b></p><p>Similar to female with the following exceptions. <i>Head</i>. Generally shorter and more compact. Clypeal process semi-elliptical, apex very broadly rounded or truncate; in profile distinctly curved. Frons punctation sparse. Ocellar triangle anterior angle weakly acute. Antenna longer, extending to posterior margin of pronotum. <i>Color</i>. All coxae and femora brown; tibiae tinged with brown medially.</p><p>MEASUREMENTS. LFW 1.07 mm; LM 0.57 mm; WH/LH 0.87; DH/LH 0.60; EV/HE 0.62; EV/WF 0.63; WF/HE 0.98; WH/WF 1.98; OOL/WOT 1.20; DM/LM 0.41; HFL/HFW 2.13; A3 L/ W 1.00; 2R1 L/ W 2.72.</p><p><b>Distribution</b></p><p>Queensland and New South Wales; known from several collections in the eastern forest zone.</p><p><b>Remarks</b></p><p>This species and <i>S. campanula</i> sp. nov. appear to form a closely related allopatric species pair, although relatively few collections are available for each. In both the clypeus is shorter than in other species of this complex, so they are keyed out twice.</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_19566073 |
| institution | Zenodo |
| language | |
| publishDate | 2026 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Sierola allithoros Magnacca, 2026, sp. nov. Magnacca, Karl Biodiversity Taxonomy Animalia Arthropoda Insecta Hymenoptera Bethylidae Sierola Sierola allithoros <p><b><i>Sierola allithoros</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: E30FF356-4328-493B-9D9C-4BE72F898135</p><p>Figs 10, 15</p><p><b>Diagnosis</b></p><p>Belonging to the complex of species with the clypeus moderately large and quadrate, visibly parallel-sided beyond the antennae. Forming a species pair with <i>S. campanula</i> sp. nov., united in having the frons very narrow (WF/HE ~0.8), clypeus with a low but distinct medial carina and broadly rounded apically rather than straight truncate or emarginate, vein Rs ending short of the anterior wing margin, and S5 with a pair of elongate setae. Separated by having the mandible reddish brown and slightly twisted, propodeum with the median carina faint, and profemur mostly yellow.</p><p><b>Etymology</b></p><p>From the Greek ‘ <i>allithoros</i> ’, ‘cross-eyed’ or ‘squinting’, from the dorsally close eyes. It is a noun in apposition.</p><p><b>Type material</b></p><p><b>Holotype</b></p><p>AUSTRALIA – <b>Queensland</b> • ♀; Upper Clayton Gully, near Cunningham’s Gap; elev. 400–600 m; 25 Apr. 1974; I. Naumann leg.; sweeping long grass; wet sclerophyll forest; QM T262151.</p><p><b>Paratypes</b></p><p>AUSTRALIA – <b>New South Wales</b> • 2 ♀♀, 1 ♂; Hat Head; 31.0626° S, 153.052° E; elev. 37 m; 4–18 Feb. 2018; P. Hebert leg.; Malaise trap, trap #3; BOLD: NSWHO3417-18, NSWHO3816-18, NSWHO3706-18, BIN ADV9054; ANIC. – <b>Queensland</b> • 1 ♀; Corroy [sic: Cooroy]; 15 Sep. 1979; G. Gordh leg.; UCRC 401812 • 1 ♀; Acacia Ridge near Brisbane; 13 Dec. 1979; G. Gordh leg.; UCRC 556776.</p><p><b>Description</b></p><p><b>Female</b></p><p>HEAD. Clypeal process semicircular, apex broadly rounded; dorsally with a distinct but narrow, sharp ridge raised slightly above the otherwise flat surface; in profile evenly sloping, nearly straight. Mandible moderately broad, more or less parallel-sided except at base; slightly twisted, outwardly shining but with distinct, fine microsculpture; in ventral view straight or nearly so. Apex of mandible slightly oblique, with teeth distinctly stronger ventrally. Malar space very small. Dorsal setae of head shorter than flagellar width. Frons with moderately close punctures between antennae and midpoint of eyes, 1–3 pit widths apart, sparse posteriorly. Vertex and gena setae similar to and only slightly longer than setae of frons. Frons microsculpture finely microreticulate to very weak and mostly smooth, shining. Vertex in dorsal view more or less straight; occipital margin broadly rounded. Lateral ocelli distinctly separate from vertex crest by less than ocellus length. Ocellar triangle anterior angle strongly acute. Head more or less parallel-sided for some distance posterior to eyes. Eye setae short, only about as long as 1–2 ommatidia. Dorsum of head in lateral view slightly convex, head deepest on line through posterior margin of eye. Venter of head posterior to mandible convex in lateral view, with relatively sparse setae. Head weakly convex ventrally, genal angle absent. Side of head posterior to eye with a line of smooth tesselate microsculpture extending from posteroventral margin of eye to vertex crest, distinctly different from smoother sculpting dorsally and ventrally. Gena ventral to eye convex; sculpture coriaceous ventrally and posterolaterally, with broad band of tesselate-colliculate sculpture extending diagonally on each side from anterior margin of eye to posteroventral margin of head. Postgenal depression absent. Antenna extending to anterior margin of pronotum.</p><p>MESOSOMA. Anterior margin of dorsal pronotal area in lateral view weakly angulate. Pronotal setae about as long as flagellar width or scarcely longer. Propodeal disc laterally coarsely rugulose; median carina faintly present; transverse carina absent, brow of declivity broadly rounded; metapleural carinae complete. Profemur strongly broadened, less than twice as long as wide, strongly laterally compressed, slightly arcuate in dorsal view.</p><p>METASOMA. Shining, apical tergites punctate laterally. S5 with a pair of long erect setae. Sixth sternum broadly glabrous medially, with sparse setae laterally.</p><p>COLOR. Body entirely dark brown to black. Antenna with scape and base of flagellum pale, becoming darker over apical half. Mandible reddish brown. Procoxa and femur yellow except dorsal margin of femur, meso- and metafemora brown, tibiae, tarsi, and trochanters yellow.</p><p>WINGS. Membrane hyaline. Basal cells of fore wing nearly glabrous, with only a few scattered setae. Cell 1M weakly convex posteriorly, corner posteroapical. Distal portion of vein Rs nearly straight but not quite meeting anterior wing margin, poststigmal abscissa of R1 indistinct.</p><p>MEASUREMENTS. LFW 1.15 mm; LM 0.62 mm (0.59–0.69); WH/LH 0.74 (0.70–0.74); DH/LH 0.52 (0.50–0.52); EV/HE 0.70 (0.64–0.76); EV/WF 0.88 (0.84–1.00); WF/HE 0.79 (0.76–0.84); WH/WF 2.18 (2.05–2.24); OOL/WOT 1.71 (1.67–1.88); DM/LM 0.40; FL/FW 1.96 (1.96–2.08); HFL/HFW 2.16 (2.00–2.33); A3 L/ W 0.83 (0.83–1.00); 2R1 L/ W 3.14. N = 5.</p><p><b>Male</b></p><p>Similar to female with the following exceptions. <i>Head</i>. Generally shorter and more compact. Clypeal process semi-elliptical, apex very broadly rounded or truncate; in profile distinctly curved. Frons punctation sparse. Ocellar triangle anterior angle weakly acute. Antenna longer, extending to posterior margin of pronotum. <i>Color</i>. All coxae and femora brown; tibiae tinged with brown medially.</p><p>MEASUREMENTS. LFW 1.07 mm; LM 0.57 mm; WH/LH 0.87; DH/LH 0.60; EV/HE 0.62; EV/WF 0.63; WF/HE 0.98; WH/WF 1.98; OOL/WOT 1.20; DM/LM 0.41; HFL/HFW 2.13; A3 L/ W 1.00; 2R1 L/ W 2.72.</p><p><b>Distribution</b></p><p>Queensland and New South Wales; known from several collections in the eastern forest zone.</p><p><b>Remarks</b></p><p>This species and <i>S. campanula</i> sp. nov. appear to form a closely related allopatric species pair, although relatively few collections are available for each. In both the clypeus is shorter than in other species of this complex, so they are keyed out twice.</p> |
| title | Sierola allithoros Magnacca, 2026, sp. nov. |
| topic | Biodiversity Taxonomy Animalia Arthropoda Insecta Hymenoptera Bethylidae Sierola Sierola allithoros |
| url | https://doi.org/10.5281/zenodo.19566073 |