Sierola claniae Magnacca, 2026, sp. nov.

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Autore principale: Magnacca, Karl
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Pubblicazione: Zenodo 2026
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author Magnacca, Karl
author_facet Magnacca, Karl
contents <p><b><i>Sierola claniae</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: 433FE20D-3EF0-433D-BAEF-FD8098E9B552</p><p>Figs 48–49</p><p><b>Diagnosis</b></p><p>Belonging to the complex of species with the clypeus moderately large and quadrate, visibly parallel-sided beyond the antennae. Distinguished from most others by having the clypeus basally raised in a narrow, convex ridge, neither carinate nor entirely flat to the base. Separated from the similar, sympatric <i>S. clavicula</i> sp. nov. by having the clypeal basomedian ridge short, apical portion of clypeus flat and polished; head narrower and more quadrate in dorsal view, parallel-sided immediately posterior of the eyes; vein Rs reaching wing margin, cell 2R1 closed; and the propodeum with a broad polished median strip from base to declivity.</p><p><b>Etymology</b></p><p>Named for the apparent host, <i>Clania lewinii</i> (Westwood, 1855) (Lepidoptera: Psychidae; recorded as <i>C. tenuis</i> Rosenstock, 1885).</p><p><b>Type material</b></p><p><b>Holotype</b></p><p>AUSTRALIA – <b>Victoria</b> • ♀; Ballarat; 9 Apr. 1954; E.F. Riek leg.; ANIC 32-154820.</p><p><b>Paratypes</b></p><p>AUSTRALIA – <b>Australian Capital Territory</b> • 1 ♀; Black Mountain; 2 Jun. 1931; L.F. Graham leg.; ANIC • 1 ♀; Canberra; 18 Jan. 1980; D.F. Rentz leg.; ex swimming pool; ANIC. – <b>New South Wales</b> • 3 ♀♀; Royalla; 21 Mar. 1961; D.H. Colless leg.; ANIC • 1 ♀; Lake George Range, 18 km N by W of Bungendore; 15 Feb. 1982; J.F. Lawrence and A. Calder leg.; on <i>Xanthorrhoea australis</i>; ANIC. – <b>South Australia</b> • 1 ♀; Mt Lofty Rgs; N.B. Tindale leg.; SAMA 32-42903. – <b>Victoria</b> • 1 ♀; 18 km NW by N of Omeo; 28 Feb. 1980; I.D. Naumann and J.C. Cardale leg.; ex alcohol collection; ANIC • 2 ♀♀; on moth <i>Clania tenuis</i>; P. Cameron leg. 1914–110; NHMUK 13379657.</p><p><b>Description</b></p><p><b>Female</b></p><p>HEAD. Clypeal process quadrate, apex broadly rounded or truncate; dorsally with a rounded carina dorsally, effaced towards apex (broadened apical portion flat); in profile evenly sloping, nearly straight. Mandible moderately narrow, slightly narrowed medially from base, ventral margin weakly concave; not twisted, at most weakly convex outwardly, outwardly polished and shining distal of bend, coriaceous and dull basally; in ventral view obtusely elbowed. Apex of mandible vertical, with equally strong teeth. Malar space 1/3 as long as width of base of mandible. Dorsal setae of head shorter than flagellar width. Frons with moderately close punctures between antennae and midpoint of eyes, 1–3 pit widths apart, sparse posteriorly. Vertex and gena setae similar to and only slightly longer than setae of frons. Frons microsculpture evident but smooth, or nearly absent, polished. Vertex in dorsal view very slightly concave; occipital margin narrowly rounded. Lateral ocelli distinctly separate from vertex crest by less than ocellus length. Ocellar triangle anterior angle weakly acute to right. Head more or less parallel-sided for some distance posterior to eyes. Eye setae short, only about as long as 1–2 ommatidia. Dorsum of head in lateral view slightly convex, head deepest on line through posterior margin of eye. Venter of head posterior to mandible convex in lateral view, with relatively sparse setae. Head evenly convex ventrally, genal angle absent. Side of head posterior to eye convex. Gena ventral to eye convex; sculpture weakly microreticulate ventrally and posterolaterally, with broad band of tesselate-colliculate sculpture extending diagonally on each side from anterior margin of eye to posteroventral margin of head. Postgenal depression shallow, rounded. Antenna extending to pronotum.</p><p>MESOSOMA. Anterior margin of dorsal pronotal area in lateral view weakly angulate. Pronotal setae short, inconspicuous. Propodeal disc laterally coarsely striate; median carina absent, median area with a broad polished strip extending from base to declivity; transverse carina absent, brow of declivity broadly rounded; metapleural carinae complete. Profemur robust, about twice as long as wide and strongly convex dorsally, strongly laterally compressed, slightly arcuate in dorsal view.</p><p>METASOMA. Highly polished, without a trace of microsculpture. S2–5 each with a single sparse apical row of setae. Sixth sternum narrowly glabrous medially, laterally with dense pilose setae.</p><p>COLOR. Body entirely dark brown to black. Antenna brown on base of scape and apical flagellomeres, yellow medially. Mandible reddish brown to orange. Coxae and femora brown, tibiae dark brown, trochanters, tarsi, and bands at bases and apices of tibiae orange.</p><p>WINGS. Membrane tinged with yellow. Basal cells of fore wing nearly glabrous, with only a few scattered setae. Cell 1M weakly convex posteriorly, corner posteroapical. Distal portion of vein Rs strongly curved, meeting anterior wing margin at oblique angle.</p><p>MEASUREMENTS. LFW 1.96 mm; LM 1.00 mm (0.76–1.03); WH/LH 0.78 (0.76–0.80); DH/LH 0.52 (0.52–0.54); EV/HE 0.65 (0.56–0.67); EV/WF 0.60 (0.57–0.64); WF/HE 1.08 (0.98–1.13); WH/WF 1.74 (1.70–1.81); OOL/WOT 1.28 (1.28–1.44); DM/LM 0.40; FL/FW 1.90 (1.83–2.04); HFL/HFW 2.38 (2.21–2.38); A3 L/ W 1.43 (1.25–1.43); 2R1 L/ W 2.67. N = 9.</p><p><b>Distribution</b></p><p>New South Wales, Victoria, Australian Capital Territory, and South Australia. Widely distributed in the southeast and relatively common.</p><p><b>Remarks</b></p><p>Cameron labelled the NHMUK specimens <i>Sierola claniae</i> but never published the name. I have kept the name to avoid confusion, though neither of those (both mounted on one card) are designated as the holotype.</p>
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institution Zenodo
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publishDate 2026
publisher Zenodo
record_format zenodo
spellingShingle Sierola claniae Magnacca, 2026, sp. nov.
Magnacca, Karl
Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Bethylidae
Sierola
Sierola claniae
<p><b><i>Sierola claniae</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: 433FE20D-3EF0-433D-BAEF-FD8098E9B552</p><p>Figs 48–49</p><p><b>Diagnosis</b></p><p>Belonging to the complex of species with the clypeus moderately large and quadrate, visibly parallel-sided beyond the antennae. Distinguished from most others by having the clypeus basally raised in a narrow, convex ridge, neither carinate nor entirely flat to the base. Separated from the similar, sympatric <i>S. clavicula</i> sp. nov. by having the clypeal basomedian ridge short, apical portion of clypeus flat and polished; head narrower and more quadrate in dorsal view, parallel-sided immediately posterior of the eyes; vein Rs reaching wing margin, cell 2R1 closed; and the propodeum with a broad polished median strip from base to declivity.</p><p><b>Etymology</b></p><p>Named for the apparent host, <i>Clania lewinii</i> (Westwood, 1855) (Lepidoptera: Psychidae; recorded as <i>C. tenuis</i> Rosenstock, 1885).</p><p><b>Type material</b></p><p><b>Holotype</b></p><p>AUSTRALIA – <b>Victoria</b> • ♀; Ballarat; 9 Apr. 1954; E.F. Riek leg.; ANIC 32-154820.</p><p><b>Paratypes</b></p><p>AUSTRALIA – <b>Australian Capital Territory</b> • 1 ♀; Black Mountain; 2 Jun. 1931; L.F. Graham leg.; ANIC • 1 ♀; Canberra; 18 Jan. 1980; D.F. Rentz leg.; ex swimming pool; ANIC. – <b>New South Wales</b> • 3 ♀♀; Royalla; 21 Mar. 1961; D.H. Colless leg.; ANIC • 1 ♀; Lake George Range, 18 km N by W of Bungendore; 15 Feb. 1982; J.F. Lawrence and A. Calder leg.; on <i>Xanthorrhoea australis</i>; ANIC. – <b>South Australia</b> • 1 ♀; Mt Lofty Rgs; N.B. Tindale leg.; SAMA 32-42903. – <b>Victoria</b> • 1 ♀; 18 km NW by N of Omeo; 28 Feb. 1980; I.D. Naumann and J.C. Cardale leg.; ex alcohol collection; ANIC • 2 ♀♀; on moth <i>Clania tenuis</i>; P. Cameron leg. 1914–110; NHMUK 13379657.</p><p><b>Description</b></p><p><b>Female</b></p><p>HEAD. Clypeal process quadrate, apex broadly rounded or truncate; dorsally with a rounded carina dorsally, effaced towards apex (broadened apical portion flat); in profile evenly sloping, nearly straight. Mandible moderately narrow, slightly narrowed medially from base, ventral margin weakly concave; not twisted, at most weakly convex outwardly, outwardly polished and shining distal of bend, coriaceous and dull basally; in ventral view obtusely elbowed. Apex of mandible vertical, with equally strong teeth. Malar space 1/3 as long as width of base of mandible. Dorsal setae of head shorter than flagellar width. Frons with moderately close punctures between antennae and midpoint of eyes, 1–3 pit widths apart, sparse posteriorly. Vertex and gena setae similar to and only slightly longer than setae of frons. Frons microsculpture evident but smooth, or nearly absent, polished. Vertex in dorsal view very slightly concave; occipital margin narrowly rounded. Lateral ocelli distinctly separate from vertex crest by less than ocellus length. Ocellar triangle anterior angle weakly acute to right. Head more or less parallel-sided for some distance posterior to eyes. Eye setae short, only about as long as 1–2 ommatidia. Dorsum of head in lateral view slightly convex, head deepest on line through posterior margin of eye. Venter of head posterior to mandible convex in lateral view, with relatively sparse setae. Head evenly convex ventrally, genal angle absent. Side of head posterior to eye convex. Gena ventral to eye convex; sculpture weakly microreticulate ventrally and posterolaterally, with broad band of tesselate-colliculate sculpture extending diagonally on each side from anterior margin of eye to posteroventral margin of head. Postgenal depression shallow, rounded. Antenna extending to pronotum.</p><p>MESOSOMA. Anterior margin of dorsal pronotal area in lateral view weakly angulate. Pronotal setae short, inconspicuous. Propodeal disc laterally coarsely striate; median carina absent, median area with a broad polished strip extending from base to declivity; transverse carina absent, brow of declivity broadly rounded; metapleural carinae complete. Profemur robust, about twice as long as wide and strongly convex dorsally, strongly laterally compressed, slightly arcuate in dorsal view.</p><p>METASOMA. Highly polished, without a trace of microsculpture. S2–5 each with a single sparse apical row of setae. Sixth sternum narrowly glabrous medially, laterally with dense pilose setae.</p><p>COLOR. Body entirely dark brown to black. Antenna brown on base of scape and apical flagellomeres, yellow medially. Mandible reddish brown to orange. Coxae and femora brown, tibiae dark brown, trochanters, tarsi, and bands at bases and apices of tibiae orange.</p><p>WINGS. Membrane tinged with yellow. Basal cells of fore wing nearly glabrous, with only a few scattered setae. Cell 1M weakly convex posteriorly, corner posteroapical. Distal portion of vein Rs strongly curved, meeting anterior wing margin at oblique angle.</p><p>MEASUREMENTS. LFW 1.96 mm; LM 1.00 mm (0.76–1.03); WH/LH 0.78 (0.76–0.80); DH/LH 0.52 (0.52–0.54); EV/HE 0.65 (0.56–0.67); EV/WF 0.60 (0.57–0.64); WF/HE 1.08 (0.98–1.13); WH/WF 1.74 (1.70–1.81); OOL/WOT 1.28 (1.28–1.44); DM/LM 0.40; FL/FW 1.90 (1.83–2.04); HFL/HFW 2.38 (2.21–2.38); A3 L/ W 1.43 (1.25–1.43); 2R1 L/ W 2.67. N = 9.</p><p><b>Distribution</b></p><p>New South Wales, Victoria, Australian Capital Territory, and South Australia. Widely distributed in the southeast and relatively common.</p><p><b>Remarks</b></p><p>Cameron labelled the NHMUK specimens <i>Sierola claniae</i> but never published the name. I have kept the name to avoid confusion, though neither of those (both mounted on one card) are designated as the holotype.</p>
title Sierola claniae Magnacca, 2026, sp. nov.
topic Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Bethylidae
Sierola
Sierola claniae
url https://doi.org/10.5281/zenodo.19566248