| _version_ | 1866902072444583936 |
|---|---|
| author | Nurinsiyah, Ayu Savitri Neiber, Marco T. Hausdorf, Bernhard |
| author_facet | Nurinsiyah, Ayu Savitri Neiber, Marco T. Hausdorf, Bernhard |
| contents | <p><b><i>Landouria petrukensis</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: 32D7E6E9-561F-4E05-95D2-66295BFD6C5E</p><p>Figs 19, 33, 50, 52; Tables 3–4</p><p><b>Diagnosis</b></p><p><i>Landouria petrukensis</i> sp. nov. is characterized by a rather high, sharply keeled shell with small scales all over the teleoconch, a penis that is divided into a long, broad, cylindrical distal part and a slightly dilated proximal part by a narrow point and a flagellum, which is broadest near its proximal end.</p><p><b>Etymology</b></p><p><i>Landouria petrukensis</i> sp. nov. is named after its occurrence in the surroundings of Petruk Cave in Karangbolong.</p><p><b>Material examined</b></p><p><b>Holotype</b></p><p>INDONESIA • holotype (det. anat.); Karangbolong, way to Petruk Cave; 7°42′ S, 109°24′ E; 78 m a.s.l.; 5 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; D = 8.7 mm, H = 5.2 mm; MZB 19203.</p><p><b>Paratypes</b></p><p>INDONESIA – <b>Central Java</b> • 6 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′17″ S, 109°23′59″ E; 78 m a.s.l.; MZB 19273 • 2 spec.; same data as for preceding; ZMH 133409 • 1 spec.; same data as for preceding; MZB 19204 • 5 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′19″ S, 109°24′02″ E; 78 m a.sl.; MZB 19265 • 4 spec.; same data as for preceding; ZMH 133417 • 4 spec.; Karangbolong, wall and surroundings of Petruk Cave; 7°42′21″ S, 109°24′03″ E; 104 m a.s.l.; MZB 19266 • 3 spec.; same data as for preceding; MZB 19267 • 5 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′18″ S, 109°24′05″ E; 138 m a.s.l.; MZB 19274 • 1 spec.; same data as for preceding; ZMH 133407 • 23 spec.; Karangbolong, Adiraja Hill, agroforestry; 7°42′48″ S, 109°24′05″ E; 168 m a.s.l.; MZB 19271 • 4 spec.; Karangbolong, Adiraja Hill, teak plantation; 7°42′50″ S, 109°24′05″ E; 166 m a.s.l.; MZB 19275 • 3 spec.; same data as for preceding; ZMH 133414.</p><p><b>Description</b></p><p>SHELL (Fig. 50; Tables 3–4). Depressed conical, with 4.75–5.5 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles, apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 24–37% of shell diameter, hardly obscured by columellar edge.</p><p>GENITALIA (Figs 19, 33; Table 4). Atrium short; penis divided by a narrowing into a long, broad cylindrical distal part and a slightly dilated proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; flagellum moderately long, broadest near its proximal end, where it rapidly tapers, slightly crenated, without node; vas deferens narrow and long; vagina short, distally dilated; oviduct longer, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.</p><p><b>Remarks</b></p><p>With regard to the presence of scales all over the teleoconch, <i>L. petrukensis</i> sp. nov. resembles <i>L. tholiformis</i> sp. nov. (Fig. 51) and <i>L. madurensis</i> sp. nov. (Fig. 57). It differs from both species in the smaller (D 8.1–9.8 mm vs 13.0– 14.4 mm in <i>L. tholiformis</i> sp. nov. and 11.4–12.0 mm in <i>L. madurensis</i> sp. nov.), more depressed shell (D/H 1.49–1.79 vs 1.77–1.88 in <i>L. tholiformis</i> sp. nov. and 1.75–2.11 in <i>L. madurensis</i> sp. nov.) with fewer whorls (4.75–5.5 vs 6–6.25 in <i>L. tholiformis</i> sp. nov. and 5.5–5.75 in <i>L. madurensis</i> sp. nov.), an on average narrower umbilicus (U/D 0.24–0.37 vs 0.32–0.40 in <i>L. tholiformis</i> sp. nov. and 0.38–0.40 in <i>L. madurensis</i> sp. nov.) and in the penis that is divided by a constriction into a long, broad, cylindrical distal part and a slightly dilated proximal part.</p><p>We could only investigate the anatomy of the holotype, from Karangbolong in southern Central Java (MZB 19203). There are several samples from other regions of Central Java, West Java and Banten with similar shells. Thus, it is possible that this species is more widely distributed, but the identity of the samples from other regions must be confirmed anatomically.</p><p><b>Distribution</b></p><p><i>Landouria petrukensis</i> sp. nov. is known from Karangbolong in southern Central Java (Fig. 52).</p> |
| format | Recurso digital |
| id | zenodo_https___doi_org_10_5281_zenodo_5587346 |
| institution | Zenodo |
| language | |
| publishDate | 2019 |
| publisher | Zenodo |
| record_format | zenodo |
| spellingShingle | Landouria petrukensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov. Nurinsiyah, Ayu Savitri Neiber, Marco T. Hausdorf, Bernhard Biodiversity Taxonomy Animalia Mollusca Gastropoda Stylommatophora Camaenidae Landouria Landouria petrukensis <p><b><i>Landouria petrukensis</i> sp. nov.</b></p><p>urn:lsid:zoobank.org:act: 32D7E6E9-561F-4E05-95D2-66295BFD6C5E</p><p>Figs 19, 33, 50, 52; Tables 3–4</p><p><b>Diagnosis</b></p><p><i>Landouria petrukensis</i> sp. nov. is characterized by a rather high, sharply keeled shell with small scales all over the teleoconch, a penis that is divided into a long, broad, cylindrical distal part and a slightly dilated proximal part by a narrow point and a flagellum, which is broadest near its proximal end.</p><p><b>Etymology</b></p><p><i>Landouria petrukensis</i> sp. nov. is named after its occurrence in the surroundings of Petruk Cave in Karangbolong.</p><p><b>Material examined</b></p><p><b>Holotype</b></p><p>INDONESIA • holotype (det. anat.); Karangbolong, way to Petruk Cave; 7°42′ S, 109°24′ E; 78 m a.s.l.; 5 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; D = 8.7 mm, H = 5.2 mm; MZB 19203.</p><p><b>Paratypes</b></p><p>INDONESIA – <b>Central Java</b> • 6 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′17″ S, 109°23′59″ E; 78 m a.s.l.; MZB 19273 • 2 spec.; same data as for preceding; ZMH 133409 • 1 spec.; same data as for preceding; MZB 19204 • 5 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′19″ S, 109°24′02″ E; 78 m a.sl.; MZB 19265 • 4 spec.; same data as for preceding; ZMH 133417 • 4 spec.; Karangbolong, wall and surroundings of Petruk Cave; 7°42′21″ S, 109°24′03″ E; 104 m a.s.l.; MZB 19266 • 3 spec.; same data as for preceding; MZB 19267 • 5 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′18″ S, 109°24′05″ E; 138 m a.s.l.; MZB 19274 • 1 spec.; same data as for preceding; ZMH 133407 • 23 spec.; Karangbolong, Adiraja Hill, agroforestry; 7°42′48″ S, 109°24′05″ E; 168 m a.s.l.; MZB 19271 • 4 spec.; Karangbolong, Adiraja Hill, teak plantation; 7°42′50″ S, 109°24′05″ E; 166 m a.s.l.; MZB 19275 • 3 spec.; same data as for preceding; ZMH 133414.</p><p><b>Description</b></p><p>SHELL (Fig. 50; Tables 3–4). Depressed conical, with 4.75–5.5 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles, apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 24–37% of shell diameter, hardly obscured by columellar edge.</p><p>GENITALIA (Figs 19, 33; Table 4). Atrium short; penis divided by a narrowing into a long, broad cylindrical distal part and a slightly dilated proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; flagellum moderately long, broadest near its proximal end, where it rapidly tapers, slightly crenated, without node; vas deferens narrow and long; vagina short, distally dilated; oviduct longer, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.</p><p><b>Remarks</b></p><p>With regard to the presence of scales all over the teleoconch, <i>L. petrukensis</i> sp. nov. resembles <i>L. tholiformis</i> sp. nov. (Fig. 51) and <i>L. madurensis</i> sp. nov. (Fig. 57). It differs from both species in the smaller (D 8.1–9.8 mm vs 13.0– 14.4 mm in <i>L. tholiformis</i> sp. nov. and 11.4–12.0 mm in <i>L. madurensis</i> sp. nov.), more depressed shell (D/H 1.49–1.79 vs 1.77–1.88 in <i>L. tholiformis</i> sp. nov. and 1.75–2.11 in <i>L. madurensis</i> sp. nov.) with fewer whorls (4.75–5.5 vs 6–6.25 in <i>L. tholiformis</i> sp. nov. and 5.5–5.75 in <i>L. madurensis</i> sp. nov.), an on average narrower umbilicus (U/D 0.24–0.37 vs 0.32–0.40 in <i>L. tholiformis</i> sp. nov. and 0.38–0.40 in <i>L. madurensis</i> sp. nov.) and in the penis that is divided by a constriction into a long, broad, cylindrical distal part and a slightly dilated proximal part.</p><p>We could only investigate the anatomy of the holotype, from Karangbolong in southern Central Java (MZB 19203). There are several samples from other regions of Central Java, West Java and Banten with similar shells. Thus, it is possible that this species is more widely distributed, but the identity of the samples from other regions must be confirmed anatomically.</p><p><b>Distribution</b></p><p><i>Landouria petrukensis</i> sp. nov. is known from Karangbolong in southern Central Java (Fig. 52).</p> |
| title | Landouria petrukensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov. |
| topic | Biodiversity Taxonomy Animalia Mollusca Gastropoda Stylommatophora Camaenidae Landouria Landouria petrukensis |
| url | https://doi.org/10.5281/zenodo.5587346 |